NOMINATION OF THE <strong>DOLOMITES</strong> FOR INSCRIPTION ON THE WORLD NATURAL HERITAGE LIST UNESCO 102 in particular) have allowed the eagle to regain a foothold in all the niches available, and the Dolomite area now hosts a few dozen bree<strong>di</strong>ng pairs. Major stu<strong>di</strong>es and projects, inclu<strong>di</strong>ng one project currently in progress (an Interreg project, with the participation of the Parco Nazionale Dolomiti Bellunesi and Parco Naturale Fanes-Senes-Braies/Fanes-Senes-Prags), have drawn up an outline of the situation and monitored a number of pairs, provi<strong>di</strong>ng results that are also of significant importance for administrative purposes (protection of nesting areas). Another important member of the birds of prey is the goshawk (Accipiter gentilis), which can be seen at elevations up to 2,500 m. The griffon vulture (Gyps fulvus) is also frequently sighted in the southern Dolomites, due to the reintroduction efforts made of the species in Friuli-Venezia Giulia. The golden eagle (Aquila chrysaetos) populates large expanses of the Dolomites in relatively stable populations. Several species are moving into the high pastures and beyond, inclu<strong>di</strong>ng the common swift (Apus apus and A. melba), the crag martin (Ptyonoprogne rupestris), the common raven (Corvus corax), the Alpine chough (Pyrrhocorax graculus), the black-capped chickadee (Parus atricapillus), the black redstart (Phoenicurus ochruros), the whinchat (Saxicola rubetra), the Alpine accentor (Prunella collaris), the brambling (Fringilla montifringilla), the snow bunting (Plectrophenax nivalis) and the whitewinged snowfinch (Montifringilla nivalis), which has been observed at elevations up to 2,500 m. The most notable species for aspect and habits is the wallcreeper (Tichodroma muraria), which is the European vertebrate most adapted to life on the craggy walls of the high mountains. Of the snakes, the most frequent and widespread is the common adder (Vipera berus), while the nose-horned viper (Vipera ammodytes) is extremely localized, with populations in the Val <strong>di</strong> Fiemme, in the valley of T. Cordevole, in the center of Cadore and in the Dolomiti Friulane. As for the lizards, the most widespread and characteristic species found on the pastures above 2,000 m in elevation is the viviparous lizard (Zootoca vivipara), which can subsist at altitudes even above 2,500 m. Still fairly unknown, due to its similarity to other species, is the <strong>di</strong>stribution of the Horvath lizard (Iberolacerta horvathi), which lives nearly exclusively on the rocky walls and in stony accumulations, up to 2,000 m in al-
NOMINATION OF THE <strong>DOLOMITES</strong> FOR INSCRIPTION ON THE WORLD NATURAL HERITAGE LIST UNESCO 103 titude. Among the amphibians, the most characteristic is the Alpine salamander (Salamandra atra) which has been observed up to 2,300 meters in the Dolomites. The same altitudes also give a home to the male Alpine newt (Triturus alpestris). Several species of invertebrates have adapted to the higher elevations in the Dolomites. Among the coleopterans, for example, the species Cychrus caraboides, Carabus creutzeri kircheri, Notiophilus biguttatus, Amara erratica, Harpalus laevipes, Trichotichnus laevicollis, Que<strong>di</strong>us alpestris, Teuchestes fossor, Athous subfuscus, Otiorhyncus prolixus, O. montivagus, O. nodosus, O. chalceus, O. hadrocerus, O. cadoricus, O. subdentatus, and O. subquadratus have been observed; among the <strong>di</strong>pterans, Bibio pomonae, Scaeva pyrastri, and Platycheirus manicatus are found up to 2,500 meters and higher. Among the daytime butterflies of the high elevation grazing land and screes, in ad<strong>di</strong>tion to the spectacular Apollo butterfly (Parnassius Apollo), which is not found at elevations beyond 1,600 m, noteworthy are its cousin, the clouded Apollo (P. mnemosyne), Pieris calli<strong>di</strong>ce, the mountain clouded yellow (Colias phicomone), the Alpine heath (Coenonympha gardetta), the silky ring- let (Erebia gorge), the eriphyle ringlet (E. eriphyle), the large ringlet (E. euryale), the blind ringlet (E. pharte), the common brassy ringlet (E. cassioides), the woodland ringlet (E. medusa), the dewy ringlet (E. pandrose), the sooty ringlet (E. Pluto), the Shepherd’s fritillary (Boloria pales), the mountain fritillary (B. napaea), the mazarine blue (Cyaniris semiargus), and the Alpine blue (Plebeius (Albulina) orbitulus). At very high elevations, up to 2,500 meters, several species of butterfly with extraor<strong>di</strong>nary ecological value fill the air, such as the lesser tortoiseshell and the painted lady (Vanessa urticae and V. cardui), on the paths of the long-range migratory movements that cross the Dolomites. The blooms of the high elevation pioneer plants are visited by several spectacular hymenopters apoidea, such as Psithyrus flavidus and Bombus elegans, which subsist at elevations above 2,500 m, and Bombus pyrenaeus, which has been sighted at over 3,000 m. Among the <strong>di</strong>pterans, well represented in the higher belts are the Chironomidae and Sciaridae, two families that on the Dolomites include two small-winged endemic species, respectively Bryophaenocla<strong>di</strong>us thaleri and Lycoriella ventrosa, both found at over 3,000 meters. Among the small-winged insects, an interesting specimen is the mecoptera Boreus hyemalis, which is active mainly in the winter and which has been observed in the Dolomites up to 3,200 meters. Among the arthropods, which have been sighted up to 2,500 m and beyond, on the Dolomites, several spiders such as Alopecosa pulverulenta, Gnaphosa ba<strong>di</strong>a, Pardosa nigra and Erigone tirolensis (the last two species have been sighted above 3,000 m), myriapods such as Glomeris helvetica (a circumalpine species with a fragmented <strong>di</strong>stributional range), Leptoiulus alemannicus and L. montivagus and isopod crustaceans such as Trachelipus ratzeburgii. Endemic in the Dolomites are the spider species Lepthyphantes merretti and L. brunneri. The terrestrial mollusc community is very extensive in the most peripheral <strong>di</strong>stricts. A recent survey conducted in the Parco delle Dolomiti Bellunesi (Dalfreddo et al. 2000) has led to classification of some 134 species, 111 which are land-inhabiting and 23 live in fresh water. Of this quota, some 75 species are found in the subalpine transitional belt between 1,600 and 1,900 m and 47 species reach or exceed 1,900 m. This population is much poorer in the more internal areas, for edaphic (the molluscs are much more scarce on non-carbonate soils), climatic and historic reasons. Also noteworthy are the Truncatellina monodon, present at higher elevations on the Sciliar, the Catinaccio, and the Alpe <strong>di</strong> Fedaia, and the rare Neostyriaca corynodes. Among the Annelida phylum, two species of Lumbricus subsist at very high elevations, despite the low depth of the soil. These are Dendrobaena rubida, up to 2,300 meters, and Lumbricus rubellus up to 2,400 meters. Among the aquatic species, the small leech Helobdella stagnalis reaches habitats up to 2,500 meters.