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The EffĘcts of In Partial 3ėĄLfil1n€nt of Tle B€qüireûents for Eh ...

The EffĘcts of In Partial 3ėĄLfil1n€nt of Tle B€qüireûents for Eh ...

The EffĘcts of In Partial 3ėĄLfil1n€nt of Tle B€qüireûents for Eh ...

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voluntary u¡skil-I€d movenênts .der€ ini-tiateú by thê<br />

associatíoa cort€x vhÍclì ¡'Je.rë first tra¡sfêrêd tÕ the<br />

motor cort€x a¡d tber throìlgà tle pyrani4al tract do1rn tÐ<br />

the spinal notoE cent€rs. f¡e finã1 cutcoûe '{as chsc.ked<br />

by ttìe assÕciaticå corter ria the seEsory paf-h¡rays, ?his<br />

systen <strong>of</strong> ccntrol is ã 'larg€ negativ€ feedback loop<br />

tt<strong>for</strong>ned Èhraugh the eaternal. sorld'r {fto,1970). During<br />

the learning phase, tb€ c€rebral cortex is continuously<br />

araf,e cf Èhe systenrs per<strong>for</strong>mance aÃd adjustment are Baðe<br />

as required. r,iith suf fic̀nt traini.ng, the perf,ormance<br />

cf the CArs b€come refiÈ€d vÕl-untary skill-ed rnovements.<br />

<strong>The</strong> large negatis€ fê€dback J,oop via lhe extêrnaL íor1ä<br />

is reFlace¡l by an intêrnal oÌìe trpassing tbrough the<br />

cerebellun lih:ch ¡rcald s€ q€ as a ûoàel <strong>of</strong> thê conbina-<br />

tion cf tbe spinal motor systên,th€ êxternal r¡orld and<br />

th+ ,sensory pêth*ays[, Yhus the oxj.gj"na]- legative feed-<br />

back system j.s corì v€rted by learai*g into a feed<strong>for</strong>d<br />

systå¡D flhlch no longer dep€nd on a stEaig¡t<strong>for</strong>d negatj-ve<br />

feedback lcop frou the output to the i¡put. Àilditional<br />

j.!ìput path{ays to th€ ce¡ebell-ì¡û s€ry€s to adjlrst t}re<br />

internal norl€Ì as requ.ir€d in a ser eonech ani sn- li ke<br />

syst êm "<br />

fn tbe contêxt ot ItÐrs a nal ysis, cerebel lar ilysn€tria<br />

a¡d int€ntionai- tr€aor rere vi-e¡,e d. as a result <strong>of</strong><br />

loss or :uFairnert <strong>of</strong> th€ irternal qodel. <strong>The</strong> iepair ê¡t<br />

Hould na!:it€st itselt in the oterall perf ori¡ance <strong>of</strong> a<br />

system {hj.cb ¡ltilized a .largê ¡egative feerlþack loop vith<br />

a ccn.p1€x t¡ansfer functicr. As is shoun in the metho(l<br />

215

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