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bishop museum bulletins in cultural and environmental studies

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Carlton & Eldredge — Mar<strong>in</strong>e Bio<strong>in</strong>vasions of Hawai‘i<br />

9<br />

Intercept: associated with fisheries products or fouled vessels brought to Hawai‘i, with the species<br />

concerned hav<strong>in</strong>g subsequent access to open ocean waters. Examples <strong>in</strong>clude the polychaete<br />

Boccardia proboscidea, the crab Charybdis helleri, <strong>and</strong> the alga Sargassum muticum.<br />

Deleted: one or a few <strong>in</strong>dividuals collected more than 50 years ago, <strong>and</strong> for which subsequent<br />

searches have not revealed the species. Examples <strong>in</strong>clude the isopod Sphaeroma terebrans,<br />

the mudshrimp Upogebia pugettensis, the crab Carc<strong>in</strong>us maenas <strong>and</strong> the sea spider<br />

Anoplodactylus eroticus.<br />

Released; Not Established: <strong>in</strong>tentional release but with no subsequent evidence of hav<strong>in</strong>g<br />

established a naturally reproduc<strong>in</strong>g population <strong>in</strong> the wild. Examples <strong>in</strong>clude the snails<br />

Haliotis spp. <strong>and</strong> Trochus niloticus, the shrimp Macrobrachium rosenbergii, <strong>and</strong> numerous<br />

mar<strong>in</strong>e fish species.<br />

Escaped; Not Established: species that escaped from culture operations, but with no subsequent<br />

evidence of hav<strong>in</strong>g established a naturally reproduc<strong>in</strong>g population <strong>in</strong> the wild. Examples<br />

<strong>in</strong>clude penaeid shrimp, the alga Mazzaella volans, <strong>and</strong> the alga Macrocystis pyrifera.<br />

CRYPTOGENIC:<br />

Species that are not clearly native or <strong>in</strong>troduced (Carlton, 1996a).<br />

NATIvE:<br />

Species present prior to human contact. endemic species are found only <strong>in</strong> Hawai‘i; <strong>in</strong>digenous<br />

species are native but not unique to the Isl<strong>and</strong>s.<br />

ACKNOWLEDGMENTS<br />

We are grateful to a large number of colleagues who have provided advice, assistance, clarification<br />

of records, repr<strong>in</strong>ts, <strong>and</strong> excavation of obscure literature. These <strong>in</strong>clude Isabella Abbott, Shane<br />

Ahyong, James Allen, Julie Bailey-Brock, Rol<strong>and</strong>o Bastida-Zavala, Michael Berrill, Rudiger Bieler,<br />

Chip Biernbaum, Barbara Butler, David Ronald Bybee, Stephen Cairns, Dale Calder, Roy Caldwell,<br />

Debby Carlton, John Chapman, Donald Ch<strong>and</strong>ler, C. Alan Child, Eugene Coan, Andrew Cohen,<br />

L<strong>in</strong>da Cole, Steven Coles, Rachel Coll<strong>in</strong>, William Cooke, Robert Cowie, Gerald L. Crow, Peter<br />

Davie, Michael Dawson, Am<strong>and</strong>a Demopoulos, Matthew Dick, Bert Draper, Ralph DeFelice,<br />

Ronald Englund, Christer Erséus, Neal Evenhuis, Daphne Faut<strong>in</strong>, Darryl Felder, Paul Fofonoff,<br />

William Font, Charles Galt, Jonathan Geller, Lisa-Ann Gershw<strong>in</strong>, Christopher Glasby, Scott<br />

Godw<strong>in</strong>, D. Goodw<strong>in</strong>, Terry Gosl<strong>in</strong>er, David Greenfield, Danièle Gu<strong>in</strong>ot, David Gulko, Eduardo<br />

Hajdu, Michael Hadfield, S.N. Harkantra, Leslie Harris, Richard Heard, Robert Hershler, Brenden<br />

Holl<strong>and</strong>, Charlotte Holmquist, Bridget Holohan, John Hooper, John Hoover, Frank Howarth, Samuel<br />

Ensung Kahng, Reg<strong>in</strong>a Kawamoto, Alison Kay, Robert K<strong>in</strong>zie, Phyllis Knight-Jones, Alan Kohn,<br />

Gretchen Lambert, Janet Lamberson, James Lowry, Josh Mackie, Antonio Frias Mart<strong>in</strong>s, Karen<br />

Maruska, George Matsumoto, L<strong>in</strong>da McCann, Claudia Mills, Anthony Montgomery, Bruce Mundy,<br />

Todd Newberry, William Newman, Peter Ng, Gordon Nishida, Arne Nygren, Daniel Otte, Brian<br />

Paavo, Gustav Paulay, Sara Pelleteri, Mary Petersen, Andrew Pitman, Cory Pittman, Richard Pyle,<br />

John E. R<strong>and</strong>all, Geoff Read, David Reid, Raul Aguilar Rosas, Gary Rosenberg, Michael Roy,<br />

Dennis Russell, Katsushi Sakai, Al Samuelson, Marilyn Schotte, Michelle Kelly-Borges Shanks,<br />

Alison Sherwood, Karla McDermid Smith, Michael Stat, Isabel Stirl<strong>in</strong>g, Stefano Taiti, C. Takahashi,<br />

Harry ten Hove, Thomas Theeger, Wesley Thorsson, Rob van Soest, Roy Tsuda, Cather<strong>in</strong>e Unabia,<br />

Daniel Wagner, Kerst<strong>in</strong> Wasson, Les Watl<strong>in</strong>g, Judith W<strong>in</strong>ston, Robert Woollacott, Brian Wysor,<br />

Chela Zab<strong>in</strong>, <strong>and</strong> Helmut Zibrowius. Fund<strong>in</strong>g for our work was provided by The Charles H. <strong>and</strong><br />

Margaret B. Edmondson Trust to which we are deeply thankful.

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