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The Application of Ooubled Haploid Plants to Population ... - MSpace

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2.7<br />

<strong>The</strong>re is potential for loss <strong>of</strong> desirable characteristics if not induded in the initial<br />

selection criteria. Mass and pedigree selection are only useful in improving B. tapa if<br />

there is enough diversity within the selected populations <strong>to</strong> avoid inbreeding<br />

depression. Boü, <strong>of</strong> these procedures require many cydes <strong>of</strong> selection <strong>to</strong> obtain a<br />

stable population with desirable characteristics<br />

Synthetic varieties are developed by crossing parents seleded based on their<br />

performance and general combining ability as determineci by the performance <strong>of</strong><br />

progeny from different combinations (Becker, 1988). <strong>The</strong> synthetic is established by<br />

mixing seed and allowing interpollination for a number <strong>of</strong> generations. Parents can be<br />

clones. inbred lines or small populations and must be maintained in isolation so that the<br />

variety can be reconstituted perbdically. <strong>The</strong> number <strong>of</strong> parents selected <strong>to</strong> create the<br />

synthetic must be large enough <strong>to</strong> minimize inbreeding depression and small enough <strong>to</strong><br />

incorporate only individuals that will maintain the mean performance <strong>of</strong> the population<br />

(Becker. 1988). <strong>The</strong> optimum number <strong>of</strong> generations <strong>of</strong> random interpollination will<br />

result in the maximum heterotygosity and potential for heterosis as shown in B. napus<br />

(Schuster, 1982).<br />

Synthetic varieties have been developed in B. mpa <strong>to</strong> explol heterosis for yield<br />

using two or more cultivars or inbred lines mixed in equal proportions and grown in<br />

isolation (Buna, 1995). Development <strong>of</strong> synthetics is not currently used in B. napus<br />

breeding because it is difficult <strong>to</strong> predict the proportion <strong>of</strong> outcrossing between the<br />

parental lines based on the genetic variation and environmentally induced variation in<br />

outcrossing rates in B. napus (Rakow and Woods. 1987). Synthetics in B. repa have

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