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Palynology of Middle Tertiary lacustrine deposits

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220 K. TAKAHASHI & U. lux<br />

Tricolpopollenites d. minutus Gonzalez Guzman n. comb.<br />

PI. 22, figs. 17-18.<br />

1966 Psilatricolpites minutus Gonzalez Guzman, Van Hoeken-Klinkenberg, Leidse Geol. Meded.,<br />

vol. 38, p. 41, pI. II, fig. 6.<br />

1967 Psilatricolpites minutus Gonzalez Guzman, B. J. Brill, Leiden, p. 27, pI. 1, Figs. 2­<br />

2 e.<br />

Diagnostic characters: Tricolpate pollen <strong>of</strong> ellipsoidal to broad-ellipsoidal or prolate to<br />

subprolate shape and rounded apices. The three narrow colpi are quite distinctly<br />

lined towards the poles. Exine chagrenate or psilate, two-layered, O. 5-1. 5 pm<br />

thick.<br />

Measurements: 21 - 24 pm X 14 - 19 pm in size.<br />

Width / length ratio: 0.66-0.79.<br />

Previous records: South America (<strong>Middle</strong> Eocene) and Africa (<strong>Middle</strong> Eocene).<br />

Colombia: <strong>Middle</strong> Eocene (Tibu area) .<br />

Nigeria: <strong>Middle</strong> Eocene (Gbekebo).<br />

Remarks: Van der Hammen (1956, p. 88) defined the subgenus Psilatricolpites with<br />

in the genus Tricolpites Erdtman 1947, he presented Tricolpites (Psilatricolpites) incomptus<br />

v. d. Hammen 1956 as subgenotype by selecting the pollen <strong>of</strong> recent Bartsia<br />

santalinaefolia (HBK) BEMTH.<br />

(Scrophulariaceae) as the type-specimen.<br />

Pierce (1961) promoted the subgenus Psilatricolpites to a separate form-genus based<br />

on Psilatricolpites incomptus (v. d. Hammen) Pierce as type species. V. d. Hammen<br />

and Wymstra (1964) proposed Psilatricolpites clarissimus (v. d. H.) as lectogenotype.<br />

Potonie 0966, p. 163) again referred to another type species: Psilatricolpites<br />

psilatus Pierce 1961. After all, Psilatricolpites Pierce is nomenc1aturally invalid and<br />

the designation <strong>of</strong> P. clarissimus and P. psilatus as type species contradicts the<br />

ICBN.<br />

Botanical affinity: Cupuliferae (?)<br />

Tricolpopollenites densus Pflug<br />

PI. 22, figs. 21-22.<br />

1953 Tricolpopollenites densus Pflug, Thomson & Pflug, Palaeontographica, B, 94, S. 96,<br />

Taf. 11, Fig. 55-58.<br />

Diagnostic characters: Tricolpate pollen. Figura spheroidal with circular to subcircular<br />

contour in subpolar view. The three conspicuous colpi are slender and gaping;

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