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Palynology of Middle Tertiary lacustrine deposits

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<strong>Palynology</strong> <strong>of</strong> <strong>Middle</strong> <strong>Tertiary</strong> <strong>lacustrine</strong> <strong>deposits</strong><br />

185<br />

Previous palynologic investigation<br />

Not much information is available at all in respect to the biostratigraphic<br />

distribution <strong>of</strong> palynomorphs in <strong>Tertiary</strong> sediments <strong>of</strong> Northern Nigeria, i. e. the<br />

los Plateau. In Niger and Mali the Paleogene section <strong>of</strong> the Iullemmeden Basin commences<br />

with clay-, marl- and limestones. Certain beds contain marine micr<strong>of</strong>ossils<br />

which refer to the Paleocene (Boudouresque et aI., 1982). During the Eocene, the<br />

facies changed drastically to rather marginal marine conditions <strong>of</strong> sedimentation, immediately<br />

noticeable by the deposition <strong>of</strong> clayey sediments (palygorskite) and ferruguneous<br />

(Fe-, Mn-oxides) beds. On top <strong>of</strong> this rests a sequence symptomatically<br />

correlated with the continental Terminal <strong>of</strong> Kilian (1931; Lang et aI., 1986).<br />

In the arenaceous and pelitic horizons <strong>of</strong> supposed Eocene age micro- and<br />

macr<strong>of</strong>ossils were encountered and in the northern part <strong>of</strong> the basin even carbonaceous<br />

and lignitic partings (Lang et aI., 1986). With these remains the status<br />

<strong>of</strong> the onset <strong>of</strong> the Continental Terminal might be substantiated stratigraphically, at<br />

least in the Iullemmeden basin (Boudouresque et aI., 1982). Among the palynomorphs<br />

Echiperiporites icacinoides Sal. -Chebold. and Cicatricosisporites dorogensis Pot.<br />

& Gel!. together with Chenolophonidites costatus, Striatopollis bellus and<br />

Bomabacacidt"tes sp. were claimed to reveal a <strong>Middle</strong> Eocene savanna, but,<br />

elsewhere, C. dorogensis is a rather reliable index-fossil <strong>of</strong> Eocene-<strong>Middle</strong> Oligocene<br />

<strong>deposits</strong> (Takahashi & lux, 1982). A tropical mangrove swamp, vaguely dated with<br />

Ecocene-Miocene appears to be reflected palynologically (Spinozonocolpites echinatus<br />

Muller) at the base <strong>of</strong> the Gwandu Formation in Nigeria (Kogbe & Sowunmi, 1975;<br />

Kogbe,<br />

1979), the latter being correlated with both the Continental Terminal<br />

(Boudouresque et aI., 1982) and the Kerri-Kerri Formation (Kogbe, 1981).<br />

Palynomorphs are not easily recovered from the clastic sequence <strong>of</strong> the Gwandu<br />

Formation, but some peaty lenses which were located in a borehole at Kaloye<br />

(lullemmeden Basin, Nigeria), yielded tricolporate and monoporate grains which<br />

perhaps derived from the vegetation <strong>of</strong> swampy, riverine and tropical forests (Kogbe<br />

& Sowunmi, 1975) : Zonocostites ramonae Germeraad et aI., Spinizonocolpites<br />

echinatus Muller, Monoporites annulatus v. d. Hammen, Retitricolporites irregularis<br />

v. d. Hammen & Wymstra, Tetracolporopollenites sp. together with unidentified<br />

pollen grains which may refer to Mauritia, Arenga, Ken tia, Uapaca and Ravenea. In<br />

spite <strong>of</strong> the fact that only 70% <strong>of</strong> this assemblage could be classified systematically a<br />

dominance <strong>of</strong> dicotyledons (50%) is revealed.<br />

Conifers appear to be missing<br />

whereas monocotyledons and pteridophytes were recognized (each group with ca.<br />

10%). From the species announced the Gwandu Formation can <strong>of</strong> course only be

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