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Terrestrial Palaeoecology and Global Change

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Chapter 7. Climate change<br />

191<br />

Growth increment (sclerite) patterns on fish scales are of a potential significance for<br />

palaeoclimatology as a source of evidence on changing growth rates related to seasonal<br />

trophic conditions (Krassilov, 1983b). Fish scales from the Mid-Cretaceous (Albian) of<br />

Crimea within the redbed domain show a regular sclerite spacing interrupted by narrow<br />

zones of constricted sclerites, but lacking distinct annual rings. Such pattern indicates<br />

only a brief period of retarded growth related to seasonal draughts (Fig. 82).<br />

In spite of their apparent utility for palaeoclimatology, the climatic constraints of laterite<br />

pedogenesis, caliche, bauxites <strong>and</strong> related products of biochemical weathering, rather<br />

than remaining constant over times, change with atmospheric CO 2<br />

levels, relief, geological<br />

substrates <strong>and</strong> vegetation. Most redbeds are volcanomictic molassoids confined to<br />

post-orogenic phases of tectonomagmatic evolution (in particular, of the Hercynian <strong>and</strong><br />

Laramid orogenies). The direct sedimentological effects of tectonomagmatic activation<br />

are topographic, creating new basinal morphologies, as well as geochemical, providing a<br />

source of metallic enrichment (in particular, by iron from mafic minerals). The indirect<br />

effects are owing to the associated sea level <strong>and</strong> climatic events (II.7.2).<br />

Acidification of surface waters <strong>and</strong> rhizosphere by volcanic substrates enhances<br />

biochemical weathering promoting redbeds <strong>and</strong> their related pedogenic lithotypes far<br />

beyond their present-day climatic ranges. With peneplanation of volcanic plateaux, the<br />

Cretaceous to Eocene bauxite deposition, traditionally taken as evidence of a tropical<br />

climate, spread into the temperate vegetation zone over northern Kazakhstan <strong>and</strong> West<br />

Fig. 82. Growth increments (sclerites) on a fish scale from the Albian of northern Crimea as a climatic<br />

indicator (Krassilov, 1983a): the absence of annual rings indicates a round-year growth, with broad zones of<br />

regular sclerites attesting to constant growth rates. The intervening zones of about 5-6 narrow irregular<br />

sclerites (arrow) reflect a retarded growth over the relatively brief (four – seven weeks) periods of adverse<br />

conditions, such as trophic lows during dry seasons.

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