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Terrestrial Palaeoecology and Global Change

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Chapter 7. Climate change<br />

207<br />

Fig. 90. The relative extent of the major biomes over the Carboniferous to Neogene: (AT) Arctic tundra, (DF)<br />

Deciduous forest, (RF) Rainforest, (TF) Boreal forest (taiga), (XS) Xeric shrubl<strong>and</strong>s to grassl<strong>and</strong>s.<br />

history (Fig. 90). The Mediterranean-type climate is likewise persistent, formerly supported<br />

by the vast extent of the Tethys seas. Its occasional excursions to high latitudes<br />

(as in the Early Eocene) correspond to minimal ice caps.<br />

VII.2. Causes <strong>and</strong> effects<br />

Climate, as a complex system, comprises a number of subsystems with their own<br />

dynamics. Climate affects, <strong>and</strong> is feedback affected by circulation of oceanic water<br />

masses, carbonate deposition, burial of organic matter, weathering of geological substrates,<br />

pedogenesis, vegetation changes, etc. In each subsystem, climatic evolution with<br />

a positive feedback is directional (humidity promotes forestation that increases humidity)<br />

while negative feedbacks generate cyclicity (CO 2<br />

greenhouse enhances biotic production<br />

that reduces atmospheric CO 2<br />

).<br />

Secular climates arise at the interference of directional <strong>and</strong> cyclic processes <strong>and</strong> are by<br />

necessity less predictable than any of these. Critical for the underst<strong>and</strong>ing of climatic cyclicity

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