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Terrestrial Palaeoecology and Global Change

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Chapter 7. Climate change<br />

251<br />

<strong>and</strong> tall trees (Mastixia of montane rainforests), the fruit–seed remains prevail over leaf<br />

remains, a taphonomy indicating shrubl<strong>and</strong> rather than rainforest.<br />

At the same time, the mammalian turnover, with a modification of the crown types in<br />

the larger herbivores over the PEB (Janis, 2000; Bowen et a., 2992), indicates a change<br />

to a more open Eocene vegetation. According to a recent palynological analysis by<br />

Harrington (2001), the vegetation change was insignificant, with the Early Eocene communities<br />

slightly more diverse <strong>and</strong> “more successional”. However, a drop of Tricolpites,<br />

a dominant palynotype of the Palaeocene nemoral biome, <strong>and</strong> a rise of palms as well as<br />

herbaceous monocots that increased by about 80% indicate a change to more open<br />

vegetation. The carbonate rhizocretions of thick red palaeosols indicate a seasonality of<br />

precipitation. In the Bighorn Basin, Wyoming (Fig. 101) the reworked palaeosols contain<br />

silicified conifer wood. A subtropical climate with pronounced seasonal draughts is inferred<br />

also for the Sparnacian of Central Europe on the basis of oxygen <strong>and</strong> strontium<br />

isotope records (Schmiz & Andereasson, 2001). The palynological, taphonomic <strong>and</strong> sedimentological<br />

data are thus complementary in suggesting a major deforestation event.<br />

In a broader phytogeographic context, the anomalous advance of thermophilic vegetation<br />

to high latitudes is a regional trans-Atlantic, rather than global, phenomenon. Chro-<br />

Fig. 101. Redbed sequence over the Palaeocene–<br />

Eocene boundary, Bighorn Basin, Wyoming<br />

(photograph taken during field trip of the<br />

International Conference on “Climate <strong>and</strong> Biota<br />

of the Early Paleocene”, Powell, 2001).

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