Vascular Plant and Vertebrate Inventory of Saguaro ... - USGS
Vascular Plant and Vertebrate Inventory of Saguaro ... - USGS
Vascular Plant and Vertebrate Inventory of Saguaro ... - USGS
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Chapter 5: Bird <strong>Inventory</strong><br />
Brian F. Powell<br />
Previous Research<br />
There has been considerable bird research at the<br />
Rincon Mountain District, but no comprehensive<br />
<strong>and</strong> well-documented inventory has been<br />
completed. Monson <strong>and</strong> Smith (1985) compiled<br />
a checklist for both districts <strong>of</strong> the park, but there<br />
is no documentation <strong>of</strong> the data used to create<br />
that list. The list includes abundance categories<br />
for each major vegetation community <strong>and</strong> this<br />
information was likely based on Gale Monson’s<br />
extensive knowledge <strong>of</strong> the distribution <strong>and</strong><br />
relative abundance <strong>of</strong> birds in similar vegetation<br />
communities in the region.<br />
A few studies have investigated<br />
songbird community composition in the Sonoran<br />
desertscrub on the west side <strong>of</strong> the district near<br />
the Cactus Forest Loop Drive (Johnson <strong>and</strong><br />
Haight 1991, see also Mannan <strong>and</strong> Bibles 1989)<br />
<strong>and</strong> in the Rincon Valley (Boal <strong>and</strong> Mannan 1996,<br />
Freiderici 1998, Powell 1999, 2004). Only two<br />
multi-species, non-raptor studies have taken place<br />
in the higher elevations <strong>of</strong> the district (Marshall<br />
1956, Short 2002) <strong>and</strong> no research has taken<br />
place in the mid-elevation areas <strong>of</strong> the district or<br />
on the east slope <strong>of</strong> the Rincon Mountains. In the<br />
1980s the park was concerned about the impact <strong>of</strong><br />
non-native cavity-nesting birds on native species<br />
<strong>and</strong> they commissioned studies to investigate<br />
this (Mannan <strong>and</strong> Bibles 1989, Kerpez <strong>and</strong> Smith<br />
1990). Because <strong>of</strong> the active fire management<br />
program, park personnel have been interested<br />
in the effects <strong>of</strong> fire on the Mexican spotted owl<br />
(Willey 1998) <strong>and</strong> songbirds (Short 2002) in the<br />
high elevation areas <strong>of</strong> the district. The park<br />
contracted for periodic raptor surveys (Felley <strong>and</strong><br />
Corman 1993, Berner <strong>and</strong> Mannan 1992, Bailey<br />
1994, Griscom 2000). Park personnel surveyed<br />
three Breeding Bird Atlas blocks within the<br />
district (Short 1996) <strong>and</strong> those results are reported<br />
in Corman <strong>and</strong> Wise Gervais (2005). The Tucson<br />
Bird Count includes three low-elevation sites in<br />
the park, including Rincon Creek (TBC 2005).<br />
Single species studies have included the elf owl<br />
(Goad <strong>and</strong> Mannan 1987, Steidl 2003), Mexican<br />
spotted owl (Willey 1997, 1998b, Anderson<br />
45<br />
<strong>and</strong> Schon 1999, Steidl <strong>and</strong> Knipps 1999), buffbreasted<br />
flycatcher (Conway <strong>and</strong> Kirkpatrick<br />
2001; they also noted other species; Kirkpatrick<br />
et al. 2006), <strong>and</strong> purple martin (Stutchbury 1991).<br />
Park personnel survey periodically for the cactus<br />
ferruginous pygmy-owl <strong>and</strong> Mexican spotted<br />
owl <strong>and</strong> park staff file annual reports to the U.S.<br />
Fish <strong>and</strong> Wildlife Service (<strong>Saguaro</strong> NP, unpubl.<br />
reports) on monitoring <strong>and</strong> relevant management<br />
activities related to these species.<br />
Methods<br />
We surveyed for birds at the Rincon Mountain<br />
District from 2001 to 2003, though most <strong>of</strong> our<br />
surveys took place in the springs <strong>of</strong> 2001 <strong>and</strong><br />
2002. We used four field methods: (1) variable<br />
circular-plot (VCP) counts for diurnal breeding<br />
<strong>and</strong> spring migrant birds, (2) nocturnal surveys<br />
for owls <strong>and</strong> nightjars (breeding season), (3) line<br />
transects for diurnal birds in the non-breeding<br />
season, <strong>and</strong> (4) incidental observations for all<br />
birds in all seasons. Although line-transect<br />
surveys were not included in the original study<br />
proposal (Davis <strong>and</strong> Halvorson 2000), we felt<br />
they were important in our effort to inventory<br />
birds at the district because many species that<br />
use the area during the fall <strong>and</strong> winter may not<br />
be present during breeding-season surveys.<br />
Nevertheless, we concentrated our primary<br />
survey effort on the breeding season because bird<br />
distribution is relatively uniform in that season<br />
due to territoriality (Bibby et al. 2002). Our<br />
survey period included peak spring migration<br />
times for most species, which added many<br />
migratory birds to our list.<br />
We sampled vegetation around most<br />
VCP stations. Vegetation structure <strong>and</strong> plant<br />
species composition are important predictors <strong>of</strong><br />
bird species richness or the presence <strong>of</strong> particular<br />
species (MacArthur <strong>and</strong> MacArthur 1961, Rice et<br />
al. 1984, Strong <strong>and</strong> Bock 1990). In this report<br />
we use these data to categorize <strong>and</strong> describe bird<br />
communities. These data will also be useful for<br />
habitat association studies (e.g., Strong <strong>and</strong> Bock<br />
1990; see Appendix H for results).