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Estimating the Water Requirements for Plants of Floodplain Wetlands

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For example, changes in surface and sub-surface water and nutrientavailability are explicit controls on algae and macrophyte growth, whilemacrophytes influence surface water availability via transpiration, andsurface water movement by <strong>the</strong>ir contribution to surface roughness.The GEM is run as a unit model embedded in <strong>the</strong> cells <strong>of</strong> a spatialmodel. Modelling <strong>the</strong> water regime includes water-balance calculationsthat partition water into three separate stores (above surface, <strong>the</strong>unsaturated soil zone, and <strong>the</strong> saturated soil zone) and hydrauliccalculations <strong>of</strong> surface water movement. The macrophyte growthmodel predicts changes in photosyn<strong>the</strong>tic carbon biomass by aproduction model limited by a multiplicative environmental controlfunction that includes light, nutrients, temperature and water. Themacrophyte model is a small part <strong>of</strong> <strong>the</strong> large and complex GEM, whichis designed <strong>for</strong> exploration <strong>of</strong> large ecosystem dynamics ra<strong>the</strong>r than <strong>the</strong>specific and accurate prediction <strong>of</strong> a single component such asvegetation. Fitz et al. (1996) do not report on any attempts to validate<strong>the</strong> models described.Finally, it should be pointed out that many complex, spatiallydistributed models have been developed: <strong>of</strong> catchment and riverhydrology; <strong>of</strong> surface and sub-surface hydraulics; and <strong>of</strong> vegetationresponse. The development, refinement and even use <strong>of</strong> such modelsare major undertakings that require considerable resources, especiallyappropriate modelling expertise. The coupling <strong>of</strong> hydrologic/hydraulicmodels adds an extra level <strong>of</strong> complexity, and while examples havebeen quoted, <strong>the</strong>re are few examples <strong>for</strong> major wetland systems, noexamples <strong>for</strong> floodplain wetlands in Australia, and all examples arebetter viewed as research investigations than wetland managementapplications.90 <strong>Estimating</strong> <strong>the</strong> <strong>Water</strong> <strong>Requirements</strong> <strong>for</strong> <strong>Plants</strong> <strong>of</strong> <strong>Floodplain</strong> <strong>Wetlands</strong>

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