Linking Restoration and Ecological Succession (Springer ... - Inecol
Linking Restoration and Ecological Succession (Springer ... - Inecol
Linking Restoration and Ecological Succession (Springer ... - Inecol
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126 Karel Prach et al.<br />
steepness of slopes can be reduced, <strong>and</strong> the drainage regime altered to achieve<br />
the required moisture conditions. When large-scale earth-moving machinery<br />
is used to change site/soil conditions, it can damage the substrate through<br />
compaction, especially if the soil is wet. Where this occurs, additional treatments<br />
such as surface ripping or drainage may be needed (Montalvo et al.<br />
2002).<br />
Chemical manipulations may also need to be done, especially where there is<br />
a deficiency or toxicity of chemical elements in the soil-forming material. In<br />
many mine wastes, for example, there is almost no organic matter, a very low<br />
nutrient supply, <strong>and</strong>, depending on parent material, a soil pH between 2 <strong>and</strong> 12.<br />
Fertilizers will often need to be added if the soil material is very deficient in<br />
major plant nutrients (Bradshaw 1983, Marrs <strong>and</strong> Bradshaw 1993). Sometimes,<br />
adding soil-forming materials such as green wastes, industrial wastes (e.g.,<br />
paper pulp wastes), or sewage sludge can improve site conditions (Greipsson<br />
2002).<br />
The aim of physical <strong>and</strong> chemical manipulations is to produce a site that<br />
is well contoured into the local l<strong>and</strong>scape <strong>and</strong> with appropriate drainage or<br />
irrigation for the target ecosystem to establish, <strong>and</strong> where the soil is suitable to<br />
allow the required species to establish <strong>and</strong> grow well.<br />
6.3.3 Biological Manipulation<br />
Adding seeds of the target species to the system is the most common biological<br />
manipulation, although it is possible to use other propagules or transplants<br />
from donor communities (Hodder <strong>and</strong> Bullock 1997, Antonsen <strong>and</strong> Olsson<br />
2005). Once introduced, steps must be taken to ensure their establishment <strong>and</strong><br />
persistence.<br />
Where seeds are added, they can be treated using germination-promoting<br />
agents to ensure a rapid germination. At the same time it is often sensible<br />
to add microorganisms such as the appropriate mycorrhizas <strong>and</strong> Rhizobium<br />
bacteria if leguminous plants are to be included in the seed mix (Greipsson<br />
2002; see Chapter 3).<br />
One major issue is the provenance of seeds <strong>and</strong> of plant material derived<br />
vegetatively. Over the last 20 years, there has been a large increase in the commercial<br />
availability of seeds of seminatural biotopes from native sources, <strong>and</strong><br />
in some places seeds of species can be obtained from very localized sources<br />
(e.g., http://www.floralocale.org). On the other h<strong>and</strong>, most commercial mixtures<br />
or transplants do not respect regionality (McKay et al. 2005). In general,<br />
when selecting material for sowing or transplanting, we advise that whenever<br />
possible, seeds <strong>and</strong> other propagules should be used from local sources. When<br />
transplanted material is to be used, material of a single clone should be avoided,<br />
in order to maximize genetic diversity. High genetic diversity may reduce the<br />
risk of extinction under fluctuating environments <strong>and</strong> allow a species to occupy<br />
more microhabitats.<br />
In some situations, succession is circumvented by importing an ecosystem<br />
from elsewhere, either completely in blocks (transplants) or in part. The most<br />
usual case is to import topsoil from elsewhere; where this is done, nutrients,<br />
seed banks, <strong>and</strong> soil microflora can be imported in one operation, which then<br />
includes both the physicochemical <strong>and</strong> biological manipulation (Vécrin <strong>and</strong><br />
Muller 2003).