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Manihot Manihotoides (Euphorbiaceae) - CNCFlora

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Systematic Position of the Genus 25<br />

rosette; stipules persistent, usually more than 1.0 cm long; inflorescence usually<br />

either pistillate or staminate, rarely of both sexes; pistillate inflorescence determinate,<br />

always 2 flowered. Sect 14. Stipulares (p. 206).<br />

4. Leaves nonlobed or very shallowly lobed, the sinus less than half the length of the<br />

lobes.<br />

26. Leaves subsessile; petioles short, less than 1.0 cm long. Sect 16. Brevipetiolatae (p. 215).<br />

26. Leaves petiolate; petioles long, more than 5.0 cm long. Sect 17. Peltatae (p. 225).<br />

Sectional Relations<br />

Figures 5A and 28A indicate the linear (as contrasted with the multidimensional)<br />

relationships of the sections of the genus <strong>Manihot</strong>. In addition, Fig 5A shows the<br />

relationship of the genera <strong>Manihot</strong> and <strong>Manihot</strong>oides. In terms of biological relationship,<br />

there are, of course, many other connections between these taxa, not indicated by this<br />

type of illustration.<br />

A critical examination of the Skylines (Figs 5, 28), and of those specific taxa<br />

delimited on the basis of poor specimens (or limited numbers of specimens), indicates<br />

that it is likely that some modification of specific taxa will be necessary when more<br />

specimens are available. However, we feel confident (on the basis of both morphological<br />

and geographical grounds) that the sectional differentiations represent stable taxonomic<br />

units.<br />

One may more readily achieve a grasp of the types of taxonomic relationship evident<br />

in the genus <strong>Manihot</strong> by careful comparison of the sectional taxa than by attempting to<br />

grasp the same concepts by examination of the individual species.<br />

1. <strong>Manihot</strong> sect <strong>Manihot</strong><br />

<strong>Manihot</strong> sect Parvibracteatae Pax subsect Utilissimae Pax in Engler, Pflanzenreich IV. 147(Heft<br />

44): 66. 1910. pro parte.<br />

<strong>Manihot</strong> sect Heterophyllae Pax subsect Cartbaginenses Pax in Engler, Pflanzenreich IV.<br />

147(Heft 44): 80. 1910. pro parte.<br />

Plantae caulinae, lignosae, infernae ad mediae frutices, folia petiolatae, penitae<br />

lobatae, inflorescentia monoecia panicula.<br />

Plants caulescent, woody, low to medium shrubs; leaves widely spaced on stem;<br />

petiolate, petiole attachment basal; lamina membranaceous, deeply lobed, lobes linear or<br />

obovate. Inflorescence a monoecious panicle; bracts and bracteoles setaceous, margins<br />

usually entire.<br />

TYPE. <strong>Manihot</strong> esculenta Crantz.<br />

DISTRIBUTION. World-wide lowland tropics, apparently native to the neotropics.<br />

1. <strong>Manihot</strong> esculenta Crantz, Institutiones Rei Herbariae; nutum naturae digestae ex<br />

habitu 1: 167. 1766; Pax in Engler, Pflanzenreich IV. 147(Heft 44): 67. 1910<br />

(recognized by Pax as a synonym of M. utilissima Pohl); Ciferri, Relaz. e Mon.<br />

Agrario-Coloniali 44: 1-59. 1938; Archiv. Bot. Forli 18: 27-33. 1942; Rogers &<br />

Fleming, Econ. Bot. 27(1): 1973.<br />

Jatropha manibot L., Sp. PI. ed. 1: 1007. 1753. Janipha manihot Humboldt, Bonpland & Kunth,<br />

Nov. Gen. et Spec. 2: 84. 1817.<br />

Jatropha dulcis J. F. Gmelin, Onom. Bot. 5: 7. 1772-1778. (Pohl cites this name as Jatropha<br />

dulcis (Gesner) Gmelin). <strong>Manihot</strong> dulcis (J. F. Gmelin) Pax in Engler, Pflanzenreich IV.<br />

147(Heft 44): 71. 1910.<br />

Jatropha mitis Rottb., Descript. Surinam. 21. 1776.<br />

Jatropha janipha Loureiro non L. (L. in Mantissa 1: 126. 1771). Fl. Cochinchinensis. 718. 1793.<br />

Jatropha stipulata Vellozo, Fl. Flum. Ic. 10: 5. t 82. 1825; Jatropha stipulata Arrabida in<br />

Steudel, Nomencl. ed. 2 1: 800. 1840. (This is the same as Jatropha stipulata Vellozo, but<br />

attributed by Steudel to Arrabida, the editor of Florae Fluminensis.) Type. Vellozo s n, n v.

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