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Full-text - Norsk entomologisk forening

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Symphyla and Pauropoda 163<br />

Table 111. Number of symphyls (S) and pauropods (p) recorded at the Sl~rstad locality. Number of<br />

soil samples in brackets.<br />

1972:<br />

21 Sept.<br />

25 Oct.<br />

6 Dec.<br />

Depth of samples in cm<br />

0-3 3-6 6-9 9-12<br />

1973:<br />

9 Feb.<br />

22 Mar.<br />

10 May<br />

17 June<br />

4 Aug.<br />

4 Sept.<br />

Many pauropods but no symphyls were found<br />

in soil samples from the uppermost 3 cm of<br />

the humus. Pauropods were also extracted<br />

from the moss layer. Al1 belonged to P. lanceolatus.<br />

The results of Symphyla and Pauropoda<br />

from the regular samples are seen in Table<br />

111. The only species found were S. sz~bnuda<br />

and P. lanceolatus. The fraction of soil cores<br />

with pauropods from each sampling day<br />

varied from O to 30 OJO, with a 17.2 OJO average<br />

for the whole sampIing period. The highest<br />

number of pauropods in one soil core was 30.<br />

The proportions of adults and the different<br />

instars of P. lanceolatus from each sampling<br />

day are seen in Table IV. Third instar to<br />

adult stages of symphyls were found throughout<br />

the year in both areas.<br />

DISCUSSION<br />

Symphyla<br />

According to Michelbacher (1949) the symphyls<br />

are found in al1 soil types, but they<br />

prefer soils with an open <strong>text</strong>ure as they are<br />

not capable of burrowing their own runways<br />

in the soil.<br />

In the present survey, the symphyls were<br />

not found in the uppermost part of the soil<br />

profile (Tables 1 and 111). In those instances<br />

where they were found between 6 and 9 cm<br />

depth at Slerstad, the raw humus layer was<br />

not more than 6 cm thick. At Nordmoen the<br />

humus horizon is much thinner, and here the<br />

symphyls are often found between 3 and 6<br />

cm depth. It appears therefore that the transition<br />

between the mineral soil and the humus<br />

is an upper border for the vertical migration<br />

of the symphyls. The only possible exceptions<br />

to this are the two symphyls found between<br />

3 and 6 cm depth at Slerstad 17 June 1973.<br />

However, it should be pointed out that the<br />

mineral soil rose a few mm in this soil slice.<br />

It is well known that symphyls are found<br />

in the deeper soil (Salt et al. 1948, Michelbacher<br />

1949, Edwards 1958). Nevertheless,<br />

the conditions which these animals encounter<br />

in the mineral soil are quite different from<br />

soils which are sually considered to be<br />

optimum for symphyls. Edwards (1958) found<br />

that sand was the least favourable of symphyls.<br />

From these observations one would<br />

expect that the symphyls, like most of the

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