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Full-text - Norsk entomologisk forening

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212 A. Lillehammer<br />

Blatjemn Diura bicaudata, Arcynopteryx compacta,<br />

Capnia atra, Amphinemura standfussi<br />

and lsoperla obscura were taken. As in the<br />

case of streams and lakes, there was a reduction<br />

of species from the Sub-Alpine belt<br />

to the Midde Alpine belt. The most common<br />

carnivore was again Diura bicaudata.<br />

Distribution westwards along presumed<br />

emigration routes<br />

Seven species are restricted to eastern parts<br />

of southern Norway where most of them<br />

occur widely (Table I). Among these are<br />

N emoura avicularis, Capnia bifrons, Capnopsis<br />

schilleri and N emoura flexuosa. Even<br />

Capnia atra does not extend too far west.<br />

Therefore three of the most likely entry<br />

routes into western Norway were investigated<br />

in detail.<br />

The lowest (ca. 900 m a.s.l.), and probably<br />

the most suitable since there is almost continuous<br />

birch woodland over the pass, is via<br />

Valdres over to Lerdal, Sogn. The other two<br />

possibilities - up Hallingdal and over to Eidfjord,<br />

and from Telemark over Haukeli to<br />

Hordaland and Rogaland - are somewhat<br />

higher and go above the tree line.<br />

Collecting in these areas showed that Capnopsis<br />

schilleri, Capnia bifrons, N emoura<br />

avicularis, Capnia atra and Nemoura flexuosa<br />

had somewhat different distributions.<br />

N emoura flexuosa had the most eastern distribution<br />

of the species and was not taken<br />

during the present investigations. Capnia bifrons<br />

and Capnopsis schilleri had the next<br />

most easterly distribution. In Valdres Capnia<br />

bifrons went furthest up (Tyinkrysset) while<br />

Capnopsis schilleri has so far only been found<br />

in 0stre Slidre. Neither species was taken in<br />

Hallingdal, while in Telemark both were<br />

taken more or less the same distance west,<br />

Capnia bifrons in Hjartdal and Capnopsis<br />

schilleri in Seljord.<br />

N emoura avicularis was collected in 0stre<br />

Slidre, Svenkerud in Hallingdal, and Seljord<br />

in Telemark. In the southern part of the<br />

country they occurred as far west as at the<br />

Byglandsfjord.<br />

Capnia atra goes furthest west of the species<br />

discussed here, although absent in the<br />

south-west of the country. Further north the<br />

species occurs in Lrerdal and at Eidfjord,<br />

right down to the fjord. In the south, the<br />

most westerly locality is Grungedal in Telemark.<br />

Discussion: southern Norway<br />

Area 1, Rogaland and South Hordaland,<br />

which contains a considerable range of environmental<br />

factors, shows very great differences<br />

in faunal associations. The coastal<br />

areas are poor in species, but those present<br />

are numerous. In area 2 the same tendency<br />

is apparent, the inner area (2.1) having the<br />

most species. This gradient from inner areas<br />

towards the coast seems to indicate an important<br />

distributional trend.<br />

Within areas 2 and 3 there are pronounced<br />

differences in the fauna. In the area of the<br />

inner Oslofjord (2.1), where there is a rich<br />

cover of deciduous trees along the streams,<br />

the Capniids dominate. The same is true of<br />

area 3.1, the Glama valley, where deciduous<br />

trees are also common along the streams. In<br />

the eastern parts of both areas 2.2 and 3.2,<br />

the Capniids are poorly represented. Therefore<br />

the Capniids as a group seem to prefer<br />

streams on rich soils with deciduous trees<br />

along the streams.<br />

The Capniids also have their main distribution<br />

area in the eastern part of south<br />

Norway. Two of them, Capnia bifrons and<br />

Capnopsis schilleri, have not been taken in<br />

western Norway, and the distribution of the<br />

other two species, Capnia atra and Capnia<br />

pygmaea, is very restricted in the west. This<br />

may be connected with the factors mentioned<br />

above. South-west Norway and the southern<br />

part (Sl'Jrlandet) differ from the rest of the<br />

country in that Capnia atra seems to be<br />

absent both in high altitude lakes and streams,<br />

and in streams in lower altitudes. In the high<br />

altitude lakes, the common herbivores are<br />

N emoura cinerea and N emurella picteti. They<br />

also occur in streams. Capnia pygmaea may<br />

be common, but the species has only been<br />

taken below the alpine belts. There also seems<br />

to be a trend of a decreasing number of<br />

species as the deciduous trees and willow<br />

bushes along the streams become absent at<br />

higher altitudes. This is most pronounced<br />

among the herbivorous species. The above<br />

trends will be compared as far as possible<br />

with those in northern Norway, and some of<br />

the trends themselves will be examined<br />

further in the zonation studies.

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