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Part 7 - UNC Herbarium

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RUSCACEAE 935<br />

A genus of about 30 species, rosette shrubs and trees, of s. United States and Mexico. References: Hess in FNA (2002a); Judd<br />

(2003)=Z; Bogler in Kubitzki (1998a).<br />

1 Leaves 3-4 (-5) mm wide; fruit 4-4.5 mm long, strongly asymmetrical; [of moist flatwoods of the FL Coastal Plain]..............<br />

...........................................................................................................................................................................[N. atopocarpa]<br />

1 Leaves 4-10 mm wide, glaucous; fruit 6.5-8 mm long, symmetrical; [of dry to dry-mesic sandhills of the GA and SC Coastal<br />

Plain]...................................................................................................................................................................... N. georgiana<br />

Nolina georgiana Michaux, Georgia Beargrass, Sandhills Lily. Cp (GA, SC): sandhills, sometimes locally common on<br />

slightly less xeric lower sandhill slopes; rare (SC Rare). Late May-June; late June-August. Nc. SC south to sc. GA. This species<br />

has been attributed to FL (Small 1933), but is not included in either Clewell (1985) or Wunderlin (1982, 1998). [= RAB, FNA,<br />

K, S, Z]<br />

Nolina atopocarpa Bartlett, Florida Beargrass. Pine flatwoods and savannas. Endemic to panhandle FL (Liberty and<br />

Franklin counties) and e. peninsular FL (St. Johns and Brevard counties). [= FNA, K, S, Z]<br />

Polygonatum P. Miller 1754 (Solomon's-seal)<br />

A genus of about 57 species, of temperate Eurasia and North America (most diverse in e. Asia). The P. biflorum complex is in<br />

need of further study. References: Ownbey (1944)=Z; Judd (2003)=Y; Utech in FNA (2002a); Eigsti (1942); Therman (1950,<br />

1953); Kawano & Iltis (1963); Conran & Tamura in Kubitzki (1998a).<br />

1 Leaves pubescent on the veins beneath; flowers 7-13 mm long............................................................................. P. pubescens<br />

1 Leaves glabrous; flowers 12-21 mm long.<br />

2 Stem slender, 1.5-5 mm in diameter; plants to 9 dm tall; lower axillary peduncles terete or nearly so, with (1-) 2-3 (-5)<br />

flowers; lowest peduncle in the axil of the (1st-) 3rd (-5th) axil; larger leaves 5.5-15 cm long, 1.2-6 cm wide; lower<br />

leaves clasping to 90 (-180) degrees .......................................................................................... P. biflorum var. biflorum<br />

2 Stem robust, 5-13 mm thick below the leaves; plants to 20 dm tall; lower axillary peduncles strongly flattened, with (2-<br />

) 3-6 (-15) flowers; lowest peduncle in the axil of the (3rd-) 4th-5th (-8th) leaf; larger leaves 9-25 cm long, 3.5-13 cm<br />

wide; lower leaves clasping to 300 degrees ........................................................................P. biflorum var. commutatum<br />

Polygonatum biflorum (Walter) Elliott var. biflorum, Small Solomon's-seal. Mt, Pd, Cp (GA, NC, SC, VA): moist to dry<br />

forests; common. April-June; August-October. CT, NY, and s. Ontario west to MI, NE, and IN, south to n. FL and s. AL. In<br />

addition to the two varieties recognized for our area, P. biflorum includes several additional varieties: var. hebetifolium R.R.<br />

Gates of panhandle FL (Apalachicola River area), var. melleum (Farwell) R. Ownbey of MI and Ontario, and var. necopinum R.<br />

Ownbey from the Black Hills of SD. The complex needs additional study. See var. commutatum for discussion of its distinction<br />

from var. biflorum. [= Z; < P. biflorum – RAB, C, FNA, W, Y; = P. biflorum – F, G, in the narrow sense; < P. biflorum var.<br />

commutatum (J.A. & J.H. Schultes) Morong – K; < P. commutatum (J.A. & J.H. Schultes) A. Dietrich – S]<br />

Polygonatum biflorum (Walter) Elliott var. commutatum (J.A. & J.H. Schultes) Morong, Large Solomon's-seal, King<br />

Solomon's-seal. Mt (GA, NC, SC?, VA), Pd (NC, VA): moist forests, roadbanks; common. May-June; September-October.<br />

NH west to s. Manitoba, south to SC, GA, LA, and TX. There has been a wide divergence of opinion regarding the merits (and<br />

practicality) of distinguishing this taxon from typical P. biflorum, and the characters considered most reliable; the two taxa may<br />

differ in chromosome number and geographical distribution; they are not, however, always readily distinguished<br />

morphologically. I prefer to recognize this taxon as a variety. See references for additional discussion. [< P. biflorum – RAB,<br />

C, FNA, W, Y; = P. canaliculatum (Muhlenberg ex Willdenow) Pursh – F, G, misapplied; < P. biflorum var. commutatum (J.A.<br />

& J.H. Schultes) Morong – K; < P. commutatum (J.A. & J.H. Schultes) A. Dietrich – S; = P. commutatum – Z]<br />

Polygonatum pubescens (Willdenow) Pursh. Mt (GA, NC, SC?, VA): moist forests, especially cove forests; common.<br />

Late April-June; August-October. S. Québec west to s. Manitoba, south to nw. SC, WV, KY, IN, WI, and IA. [= RAB, C, F,<br />

FNA, G, K, W, Y, Z; = P. biflorum – S, misapplied]<br />

SCHEUCHZERIACEAE F. Rudolphi 1830 (Scheuchzeria Family)<br />

A monotypic family, circumboreal in arctic and cold temperate regions. References: Nienaber in FNA (2000); Haynes, Les, &<br />

Holm-Nielsen in Kubitzki (1998b).<br />

Scheuchzeria Linnaeus (Scheuchzeria, Pod-grass)<br />

A monotypic genus, circumboreal in arctic and cold temperate regions. References: Nienaber in FNA (2000); Haynes, Les, &<br />

Holm-Nielsen in Kubitzki (1998b).

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