Part 7 - UNC Herbarium
Part 7 - UNC Herbarium
Part 7 - UNC Herbarium
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STEMONACEAE 939<br />
TOFIELDIACEAE Takhtajan 1994 (False-asphodel Family)<br />
A family of 5 genera and about 30 species, of disjunct distribution in north temperate and subarctic areas, and in the Guayana<br />
Shield and northern Andes areas of n. South America. Reveal & Zomlefer (1998) place the Tofieldiaceae in the monotypic order<br />
Tofieldiales, only distantly related to the Liliaceae. Tamura in Kubitzki (1998a) treats this group as subfamily Tofieldioideae of<br />
the Nartheciaceae. References: Zomlefer (1997c, 1999); Tamura in Kubitzki (1998a).<br />
1 Inflorescence 1-flowered; tepals yellow; seeds yellowish; [endemic to Panhandle FL] ........................... [Harperocallis flava]<br />
1 Inflorescence a raceme or thyrse; tepals white to pale cream (fading to yellowish on dried specimens); seeds brown;<br />
[collectively widespread].<br />
2 Bracts of the inflorescence large, spathelike, acuminate-aristate at the tip; tepals 9-17 mm long; stamens (6-) 9 (-12) .....<br />
................................................................................................................................................................................... Pleea<br />
2 Bracts of the inflorescence minute; tepals 2.5-5 mm long; stamens 6.<br />
3 Inflorescence a raceme (the flower pedicels attached to the scape singly); scape smooth; flowering (late August-)<br />
late September-October.................................................................................................................................Tofieldia<br />
3 Inflorecence a thyrse (flower pedicels attached to the scape in groups of 3-7); scape scurfy-scabrous; flowering<br />
June-August ..................................................................................................................................................Triantha<br />
Harperocallis McDaniel (Harper's Beauty)<br />
A monotypic genus, perennial, of southeastern United States. References: McDaniel (1968)=Y; Zomlefer (1997c)=Z; Utech &<br />
Anderson in FNA (2002a).<br />
Harperocallis flava McDaniel, Harper's Beauty. Pineland bogs. Endemic to FL Panhandle (Franklin and Liberty counties).<br />
[= FNA, K, Y, Z]<br />
Pleea Michaux 1803 (Rush-featherling)<br />
A monotypic genus, of se. North America, sometimes included in Tofieldia. References: Zomlefer (1997c)=Z; Tamura in<br />
Kubitzki (1998a); Packer in FNA (2002a).<br />
Pleea tenuifolia Michaux, Rush-featherling. Cp (GA?, NC, SC): locally abundant in wet savannas, pocosin margins,<br />
usually in peaty soil, locally abundant in a few counties in se. NC, rare inland (very rarely as far as Cumberland County, NC);<br />
uncommon (SC Rare). September-October; October-November. A Southeastern Coastal Plain endemic: se. NC and ne. SC<br />
south to sw. GA, n. FL and s. AL, but apparently absent from s. SC and ne. GA. When in flower in wet savannas and powerline<br />
rights-of-way in Brunswick County, Pleea visually dominates areas up to hundreds of hectares. In sterile condition, it is<br />
recognizable by its leathery equitant leaves, bright red at their bases. [= RAB, FNA, GW, K, S; = Tofieldia tenuifolia (Michaux)<br />
Utech – Z]<br />
Tofieldia Hudson 1778 (Bog Asphodel)<br />
A genus of about 7-8 species, of temperate to subarctic North America and Eurasia. There is controversy about the<br />
circumscription of Tofieldia.relative to the related genera Pleea and Triantha (here recognized, but sometimes subsumed into<br />
Tofieldia. Some believe that Tofieldia, Triantha, and Pleea should be treated together in a broadly circumscribed Tofieldia<br />
(Utech 1978, Zomlefer 1997c); others that all three should be treated separately (Ambrose 1980; Packer 1993; Cruden 1991).<br />
Packer in FNA (2002a) has recently recognized Triantha, Pleea, and Tofieldia as separate genera, a conclusion followed here in<br />
part because of the ancient, relictual nature of these units. References: Zomlefer (1997c)=Z; Packer (1993); Ambrose (1980);<br />
Utech (1978); Hitchcock (1944)=Y; Tamura in Kubitzki (1998a); Packer in FNA (2002a); Cruden (1991).<br />
Identification notes: In sterile condition, Tofieldia glabra can be distinguished from Iris verna by its minutely upwardlyscabrous<br />
margins (Iris has smooth margins).<br />
Tofieldia glabra Nuttall, Carolina Bog Asphodel, White Asphodel. Cp (NC, SC): savanna-pocosin ecotones, wet savannas,<br />
seepage bogs; rare (US Species of Concern, NC Rare, SC Rare). (Late August-) late September-October; October-November.<br />
Endemic to the coastal plain and sandhills of NC and northern SC (the reports from GA are dubious). [= RAB, FNA, GW, K, S,<br />
Z]<br />
Triantha (Nuttall) Baker