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Part 7 - UNC Herbarium

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POACEAE 874<br />

F, G, HC; > P. villosissimum var. pseudopubescens (Nash) Fernald – F, G; = D. ovale ssp. pseudopubescens (Nash) Freckmann<br />

& Lelong – FNA; > P. commonsianum – HC, S; P. pseudopubescens Nash – HC, S; P. addisonii Nash – HC, S; P.<br />

wilmingtonense Ashe – HC, S; = P. ovale Elliott var. pseudopubescens (Nash) Lelong – X]<br />

Dichanthelium ovale (Elliott) Gould & Clark var. ovale, Oval-flowered Witch Grass. Cp (GA, NC, SC, VA): dry to damp<br />

sandy pinelands; rare (NC Watch List, VA Rare List). May-October. NY to WI, south to FL and e. TX. Infrequent over most of<br />

its range except FL. In our region, the D. ovale double ligule character is more evident in var. ovale, with var. addisonii often<br />

having only a single ligule about 1 mm long. Also see note under D. consanguineum, and note at end of descriptions regarding<br />

Panicum malacon (placed in synonymy here). [= K, Z; = Panicum ovale Elliott – RAB, X; = D. ovale ssp. ovale – FNA; > P.<br />

ovale – HC, S; > P. malacon Nash – HC, S; = P. ovale var. ovale – X]<br />

Dichanthelium polyanthes (Schultes) Mohlenbrock, Small-fruited Witch Grass. Pd, Mt, Cp (GA, NC, SC, VA): damp to<br />

dry soil of open woods and ditches; common (absent from the outer Coastal Plain south of the Neuse River). June-October. VA<br />

to s. IL, south to GA and e. TX. [= FNA; = Panicum polyanthes Schultes – RAB, C, F, G, GW, HC, S; = D. sphaerocarpon<br />

(Elliott) Gould var. isophyllum (Scribner) Gould & Clark – K, Z]<br />

Dichanthelium portoricense (Desvaux ex Hamilton) B.F. Hansen & Wunderlin, Puerto Rican Witch Grass. Cp (GA, NC,<br />

SC): moist pine savannas and flatwoods; frequent in North Carolina, uncommon in South Carolina. May-September. NC south<br />

to FL, west to TX, also in West Indies. [= Panicum portoricense Desvaux ex Hamilton – RAB, HC, S; = D. portoricense ssp.<br />

portoricense – FNA; < D. sabulorum (Lamarck) Gould & Clark var. thinium (Hitchcock & Chase) Gould & Clark – K, Z; = P.<br />

portoricense var. portoricense – X]<br />

Dichanthelium ravenelii (Scribner & Merrill) Gould, Ravenel's Witch Grass. Cp, Pd (GA, NC, SC, VA): dry sandy or<br />

rocky thin woods and openings, sometimes in moist soils; common (VA Watch List). May-October. DE south to FL, west to e.<br />

TX, north to IA. [= FNA, K, Z; = Panicum ravenelii Scribner & Merrill – RAB, C, F, G, HC, S]<br />

Dichanthelium scabriusculum (Elliott) Gould & Clark, Tall Swamp Witch Grass. Cp, Pd (GA, NC, SC, VA): moist, low,<br />

open or shaded woodlands, often along streams or ditches; common in Coastal Plain, uncommon in Piedmont (VA Watch List).<br />

May-October. Se. MA south to FL, west to e. TX and AR. [< Panicum scabriusculum – RAB, C, GW; > P. scabriusculum – F,<br />

HC, S; > P. aculeatum Hitchcock & Chase – F, G, HC, S; < D. scabriusculum – FNA, K, Z; > P. scabriusculum var.<br />

scabriusculum – G]<br />

Dichanthelium scoparium (Lamarck) Gould, Velvet Witch Grass. Cp, Pd, Mt (GA, NC, SC, VA): moist sandy soil of<br />

woodland openings and ditches; common (uncommon in Mountains). May-October. MA and MI south to FL and TX, also in<br />

Mexico and West Indies. The dense, velvety pubescence of the internodes, sheaths, and blades of this taxon, combined with the<br />

viscid band below the nodes, are diagnostic. [= FNA, K, Z; = Panicum scoparium Lamarck – RAB, C, F, G, GW, HC, S]<br />

Dichanthelium species 2 (=webberianum), Webber's Witch Grass. Cp (GA, NC, SC): moist pine savannas and flatwoods;<br />

uncommon. May-August. Disjunct in se. NC and SC from GA and FL. [= Panicum webberianum Nash – RAB, HC, S; < D.<br />

portoricense (Desvaux ex Hamilton) B.F. Hansen & Wunderlin ssp. patulum (Scribner & Merrill) Freckmann & Lelong – FNA;<br />

< D. sabulorum (Lamarck) Gould & Clark var. patulum (Scribner & Merrill) Gould & Clark – K, Z; < P. portoricense Desvaux<br />

ex Hamilton var. nashianum (Scribner) Lelong – X]<br />

Dichanthelium species 3 (=lancearium), Nash's Witch Grass. Cp, Pd (GA, NC, SC, VA): moist pine savannas and<br />

flatwoods, moist to dry openings in maritime forests, dry pine and oak sandhills; common in Coastal Plain, uncommon in<br />

Piedmont (VA Watch List). May-September. Se. VA to FL, west to e. TX, also in West Indies and Central America. This and<br />

D. portoricense appear to intergrade in our region, and =lancearium is treated as part of D. portoricense ssp. patulum in FNA,<br />

whose concept of ssp. patulum includes Panicum webberianum (D. species 2 here) and P. patentifolium, both of which appear to<br />

merit recognition (=webberianum as a species and =patentifolium as at least a variety). [= Panicum lancearium Trinius – RAB,<br />

C, G; > P. lancearium var. lancearium – F; > P. lancearium var. patulum (Scribner & Merrill) Fernald – F; < D. portoricense<br />

(Desvaux ex Hamilton) B.F. Hansen & Wunderlin ssp. patulum (Scribner & Merrill) Freckmann & Lelong – FNA; > P.<br />

lancearium – HC, S; > P. patulum (Scribner & Merrill) Hitchcock – HC, S; < D. sabulorum (Lamarck) Gould & Clark var.<br />

patulum (Scribner & Merrill) Gould & Clark – K, Z; < P. portoricense Desvaux ex Hamilton var. nashianum (Scribner) Lelong –<br />

X]<br />

Dichanthelium species 5 (=neuranthum), Nerved Witch Grass. Cp (GA, NC): maritime wet grasslands and wet savannas<br />

near the coast; rare (GA Special Concern, NC Significantly Rare). May-September. Disjunctly in se. NC, se. SC, GA, FL, s.<br />

MS, TX, Bahamas, Cuba, and Belize. Treated as D. aciculare ssp. neuranthum in FNA, but distinctive morphology and habitat<br />

argue for recognition as a species. Can occur with the similar-appearing D. caerulescens, from which it differs by having<br />

spikelets that are longer (1.8-2.0 mm vs. 1.4-1.8), rounded vs. obtuse to sub-acute, and pubescent vs. glabrous; longer first<br />

glumes (0.8-1.0 mm vs. 0.3-0.8); leaves 15× or more as long as wide vs. 10-15×; and a nearly strict panicle. FNA gives a<br />

spikelet length of 2-2.8 mm, well beyond the length of spikelets on specimens we have seen throughout the range. As FNA<br />

provides no synonymy, it is possible that its concept of "neuranthum" includes entities treated separately here, or entities outside<br />

the range of this flora. [= Panicum neuranthum Grisebach – RAB, HC, S; = D. aciculare (Desvaux ex Poiret) Gould & Clark<br />

ssp. neuranthum (Grisebach) Freckmann & Lelong – FNA; < D. aciculare – K, Z]<br />

Dichanthelium species 9 (=cryptanthum), Hidden-flowered Witch Grass. Cp (NC, SC): wet meadows and ditches,<br />

streamside openings (NC Significantly Rare). May-September. NC (or NJ?) to MS (or TX?) (previous concepts of this taxon<br />

and its range are unclear). In the field, this taxon can be mistaken for D. yadkinense; it is readily distinguished by its scabrous<br />

peduncle and membranous ligules. [= Panicum cryptanthum Ashe – F, HC, S; < P. scabriusculum Elliott – RAB, C, GW; < D.<br />

scabriusculum (Elliott) Gould & Clark – FNA, K, Z; = P. scabriusculum var. cryptanthum (Ashe) Gleason – G]<br />

Dichanthelium species 10 (=curtifolium), Short-leaved Witch Grass. Cp (SC), Mt (NC): bogs, sphagnous streamhead<br />

swamps, mountain streams; uncommon in Coastal Plain, rare in Piedmont and Mountains. April-September. Ranging disjunctly<br />

in w. NC and e. TN, e. SC, FL, and MS. The combination of characters is quite distinctive for the genus in our region. [=

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