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Part 7 - UNC Herbarium

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POACEAE 887<br />

sense (misapplied as to our material); < F. ovina var. ovina – F, G, HC; < F. ovina var. duriuscula (Linnaeus) W.D.J. Kock – F,<br />

G, HC, misapplied as to our material]<br />

* Festuca thurberi Vasey. Introduced in SC. [= K] {not keyed; investigate}<br />

Festuca versuta Beal, Texas Fescue. Native, east to TN. [= K] {not keyed; investigate}<br />

Glyceria R. Brown (Mannagrass)<br />

(also see Torreyochloa)<br />

A genus of about 40 species, nearly cosmopolitan. References: Tucker (1996)=Z.<br />

1 Spikelets (10-) 15-40 mm long, linear, subterete, 5-15× as long as wide; [section Glyceria].<br />

2 Lemma (6-) 7-8.5 (-10) mm long, acute to acuminate; palea longer than the lemma, extending 1.5-3 mm beyond the<br />

lemma apex.....................................................................................................................................................G. acutiflora<br />

2 Lemma 2.5-5.3 mm long, obtuse to notched; palea about as long as the lemma (ranging from shorter than the lemma<br />

and included, to projecting up to 1 mm beyond the lemma apex).<br />

3 Anthers 0.5-0.8 mm long; lemma mostly 2.5-3.5 mm long................................................................... G. arkansana<br />

3 Anthers 1.0-2.0 mm long; lemma mostly 3.6-5.5 mm long.............................................................G. septentrionalis<br />

1 Spikelets 2.5-8 mm long, ovate to oblong, 1.5-3× as long as wide; [section Hydropoa].<br />

4 Inflorescence compact (at maturity), the branches stiffly ascending to appressed, the tips never nodding; ligule < 1 mm<br />

long.<br />

5 Inflorescence branches elongate, appressed; lower internodes of the inflorescence 2-8 cm long; spikelets with 3-4<br />

flowers, 3.5-4 mm long; lemma 1.9-2.8 mm long; leaves 2-5 mm wide; [of the Mountains, rarely elsewhere] .........<br />

................................................................................................................................................................ G. melicaria<br />

5 Inflorescence branches short, stiffly ascending; lower internodes of the inflorescence 0.8-2.0 (-2.5) cm long;<br />

spikelets with 4-7 flowers, 4-8 mm long; lemma 3.0-3.7 mm long; leaves 3-10 mm wide; [of the Coastal Plain,<br />

rarely disjunct inland to the Mountains of VA]............................................................................................G. obtusa<br />

4 Inflorescence lax and diffuse (at maturity), the branches spreading to somewhat ascending, the tips often nodding or<br />

drooping; ligule 1-6 mm long.<br />

6 Veins of the lemma visible, but not raised; lemma 2.3-4.0 mm long; ligule 2-6 mm long.<br />

7 Lemma 2.9-4.0 mm long, projecting conspicuously beyond the palea; spikelets 4-8 mm long, with 5-10<br />

flowers...........................................................................................................................................G. canadensis<br />

7 Lemma 2.3-2.9 mm long, more-or-less equal to the palea; spikelets 3-5 mm long, with (2-) 3-5 (-6) flowers ...<br />

................................................................................................................................................................ G. laxa<br />

6 Veins of the lemma prominently raised; lemma 1.4-3.0 mm long; ligule 1-4 mm long.<br />

8 Lemma 1.4-2.1 mm long, usually green; first glume 0.5-1.0 mm long; second glume 0.8-1.3 mm long; culms<br />

mostly 5-12 dm tall; leaves 2-5 (-8) mm wide ...................................................................G. striata var. striata<br />

8 Lemma 2.5-3.0 mm long, purple or green; first glume 1.2-1.9 mm long; second glume 1.5-2.4 mm long;<br />

culms mostly 10-20 dm tall; leaves 5-12 mm wide.<br />

9 Glumes obtuse, subequal (the first 1.2-1.9 mm long, the second 1.5-2.4 mm long); spikelets 4-6.5 mm<br />

long, (4-) 5-9-flowered; lemmas purplish, contrasting with the pale glumes; ligule 2-4 (-5) mm long; [of<br />

various montane wetlands of VA and possibly NC]...............................................G. grandis var. grandis<br />

9 Glumes acute, unequal (the first 1.4-1.6 mm long, the second 1.8-2.1 mm long); spikelets mostly 3-4<br />

mm long, 2-4-flowered; lemmas dark green (or purplish?); ligule 1.5-3 mm long; [of high elevation<br />

seepages in the Great Smoky Mountains of NC and TN]........................................................ G. nubigena<br />

Glyceria acutiflora Torrey. Mt (GA, VA): shallow water and wet mucky soils in mountain ponds, wet pastures; rare (GA<br />

Special Concern). June-July. ME west to MI, south to DE, VA, nw. GA (Jones & Coile 1988), e. TN, and MO; also in e. Asia.<br />

[= C, F, G, GW, HC, K, W, Z; = Panicularia acutiflora (Torrey) Kuntze – S]<br />

Glyceria arkansana Fernald, Arkansas Mannagrass. Cp (VA): swamps; rare. May-June. IL south to LA and AR; disjunct<br />

in se. VA and w. NY. The appropriate treatment of this taxon needs further investigation. [= F, HC, K, Z; < G. septentrionalis –<br />

C, G; = G. septentrionalis A.S. Hitchcock var. arkansana (Fernald) Steyermark & Kučera]<br />

Glyceria canadensis (Michaux) Trinius, Rattlesnake Mannagrass. Mt (NC, VA), Cp (VA): bogs, seepages, and wet<br />

meadows; rare. June-July. Newfoundland west to MN, south to NJ, VA, nw. NC, and IL. [= C, F, G, K, Z; = G. canadensis var.<br />

canadensis – HC, W]<br />

Glyceria grandis S. Watson var. grandis, American Mannagrass. Mt (NC?, VA), Cp (VA): wet, mucky soils of open<br />

wetlands; rare. Nova Scotia west to AK, south to VA, IA, NM, and OR. Attributed to w. NC by Tucker (1996). [= K; < G.<br />

grandis – C, F, G, GW, HC, Z; = Panicularia grandis (S. Watson) Nash – S; ? G. maxima (Hartman) Holmberg ssp. grandis (S.<br />

Watson) Hultén]<br />

Glyceria laxa (Scribner) Scribner, Lax Mannagrass. Mt (NC, VA): bogs; rare. June-July. Prince Edward Island south to<br />

NC, mostly Appalachian. Though often described as a hybrid of G. canadensis and either G. striata var. striata and/or G.<br />

grandis var. grandis, G. laxa ranges south of the distribution of both G. canadensis and G. grandis var. grandis. It is best

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