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Part 7 - UNC Herbarium

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PONTEDERIACEAE 928<br />

A genus of 7-8 species, native of tropical America and Africa, but now introduced widely in warm regions. References: Cook in<br />

Kubitzki (1998b); Horn in FNA (2002a).<br />

* Eichhornia crassipes (Martius) Solms-Laub, Water Hyacinth. Cp (GA, NC, SC, VA), Pd (GA, NC): ponds, ditches,<br />

sluggish water; uncommon, introduced from South America. June-September. E. crassipes is "generally considered the world's<br />

most serious aquatic weed" (Rosatti 1987). Originally native to tropical South America, E. crassipes is now a widespread<br />

naturalized weed throughout the tropics and subtropics. In the northern part our area, water hyacinth is rare, probably not long<br />

persisting. Further south, it can be an aggressive aquatic weed. [= RAB, C, F, FNA, G, GW, K; = Piaropus crassipes (Martius)<br />

Rafinesque – S]<br />

Heteranthera Ruiz & Pavón 1794 (Mud-plantain)<br />

(also see Zosterella)<br />

A genus of 10-12 species, of tropical and temperate America and tropical Africa. References: Cook in Kubitzki (1998b); Horn<br />

(1998)=Z; Horn in FNA (2002a). Key based in part on FNA.<br />

1 Leaves narrowly linear, 20-50× as long as wide, the base attenuate; flowers solitary, the corolla yellow; stamens and anthers<br />

all alike ......................................................................................................................................................................... H. dubia<br />

1 Leaves reniform, 0.5-1.5× as long as wide, the base cordate; flowers 1-several, the corolla white or pale blue; stamens and<br />

anthers dimorphic.<br />

2 Spathe with 2-several flowers; perianth tube 3-12 mm long.<br />

3 Anthers and filaments with dark purple hairs; internode below the spathe < 1 cm long; spike with (3-) 7-16<br />

flowers, typically elongating well out of the spathe...............................................................................H. multiflora<br />

3 Anthers and filaments with white hairs; internode below the spathe > 1 cm long; spike with 2-8 flowers, typically<br />

mostly included within the spathe......................................................................................................... H. reniformis<br />

2 Spathe with solitary flower; perianth tube 11-45 mm long.<br />

4 Vegetative stems elongating only in water deeper than 5 cm; blades of petiolate leaves oblong to ovate, the base<br />

truncate to cuneate; perianth tube 15-45 mm long ....................................................................................[H. limosa]<br />

4 Vegetative stems commonly elongating; blades of petiolate leaves round to oblong, the base cordate to truncate;<br />

perianth tube 11-29 mm long ........................................................................................................... [H. rotundifolia]<br />

Heteranthera dubia (Jacquin) MacMillan, Water Stargrass. Mt (NC, VA), Pd (VA), Cp (VA): streams, rivers; uncommon<br />

(rare in NC). August-September. Québec west to WA, south to Cuba and Central America, but rare or absent in much of the se.<br />

United States. The attribution of this species to SC is in error (as by Kartesz 1999), based on a misidentified specimen (C. Horn,<br />

pers. comm.). [= RAB, F, FNA, GW, K, W; = Zosterella dubia (Jacquin) Small – C, G, S]<br />

Heteranthera multiflora (Grisebach) Horn. Cp (NC, VA): in shallow, stagnant water in floodplains, or emersed on mud;<br />

rare (NC Watch List). June-October. IL west to NE, south to MS; also on the Atlantic Coastal Plain from NJ south through PA<br />

to ne. NC; also in South America (Brazil, Paraguay, Argentina, and Venezuela). [= C, FNA, K, Z]<br />

Heteranthera reniformis Ruíz & Pavón. Cp (NC, VA), Pd (GA, NC, SC, VA): in shallow, stagnant water in floodplains,<br />

or emersed on mud; uncommon (SC Rare). June-October. CT west to NE, south to FL and TX and into South America. First<br />

reported for South Carolina by Hill & Horn (1997). [= RAB, C, F, FNA, G, GW, K, S, W, Z]<br />

Heteranthera limosa (Swartz) Willdenow occurs east to TN, KY, and FL (Kartesz 1999); it is attributed to VA in Small<br />

(1933), but the documentation is not known. [= FNA, C, F, G, K, S, Z]<br />

Heteranthera rotundifolia (Kunth) Grisebach. Ponds. Midwestern, as a rare disjunct east to c. KY (Larue County) (Medley<br />

1993). [= FNA, C, K, Z]<br />

Pontederia Linnaeus 1753 (Pickerelweed)<br />

A genus of 3-6 species, from North America to South America. References: Lowden (1973)=Z; Cook in Kubitzki (1998b); Horn<br />

in FNA (2002a).<br />

1 Floral tube villous when young, essentially glabrous to sparsely glandular in maturity; leaves primarily ovate to triangularlanceolate,<br />

2.2-21 cm wide, the base generally cordate or truncate (rarely cuneate).............................. P. cordata var. cordata<br />

1 Floral tube persistently pubescent with short glandular hairs; leaves lanceolate, 0.4-8.3 cm wide, the base generally cuneate<br />

to truncate ..........................................................................................................................................P. cordata var. lancifolia<br />

Pontederia cordata Linnaeus var. cordata, Heartleaf Pickerelweed. Cp, Pd (GA, NC, SC, VA), Mt (NC): marshes, pondshores,<br />

lake-shores; common, uncommon in Piedmont, rare in Mountains. May-October. Nova Scotia west to MN, south to FL<br />

and TX; Belize; s. Brazil, Argentina, Paraguay, and Uruguay. The recognition of infraspecific taxa in Pontederia cordata is<br />

controversial and requires additional study. P. cordata exhibits tristyly, an interesting breeding system. Each plant has one of 3

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