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Part 7 - UNC Herbarium

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POACEAE 900<br />

per node, loosely flowered; spikelets 4.7-7.7 mm long; first glumes 2.5-5.5 mm<br />

long, 2/3-3/4 as long as the spikelet, 7-9 nerved, the nerves thickened and<br />

raised; fertile lemma 1.3-1.8 mm wide ........................ P. amarum var. amarum<br />

22 Rhizomes usually short; culms usually tufted, 1-2 (-3) m tall; leaves 2-5 dm<br />

long; panicles 3-10 cm wide, the primary branches usually 2 or more per node,<br />

densely flowered; spikelets 4.0-5.9 mm llong; first glumes 2-3.5 mm long, ½-<br />

2/3 as long as the spikelet, 3-5 (-7) nerved, the nerves thin and wiry; fertile<br />

lemma 1.0-1.5 mm wide ......................................................................................<br />

..................................................................................P. amarum var. amarulum<br />

21 Panicle with divergent to spreading-ascending branches; upper sheaths shorter than<br />

internodes; spikelets 2.8-5 mm long; fertile lemma 2-2.6 mm long.<br />

23 Spikelets 2.8-3.5 mm long; first glume 1/2 length of spikelet, blunt to acute ......<br />

.....................................................................................P. virgatum var. cubense<br />

23 Spikelets 3.5-5 mm long; first glume 2/3 length of spikelet, acuminate ..............<br />

................................................................................... P. virgatum var. virgatum<br />

17 Plants with hard crowns, lacking rhizomes; fertile lemma 1.2-1.6 mm long; [subgenus Agrostoides,<br />

section Agrostoidea].<br />

24 Ligule of white hairs 0.5-3 mm long; culms to 1 m long; cauline blades 2-8 mm wide, usually<br />

pilose adaxially near the base; spikelets 2.0-4.0 mm long.<br />

25 Ligules 1-3 mm long; spikelets 2.0-2.7 mm long, 2.5-4× as long as wide, often obliquely<br />

set on the pedicels..............................................................P. longifolium var. longifolium<br />

25 Ligules 0.5-1.5 mm long; spikelets 2.4-4.0 mm long, 3.5-5× as long as wide, erect on the<br />

pedicels.................................................................................... P. longifolium var. combsii<br />

24 Ligule a tawny membrane 0.5-1.0 mm long, often erose or lacerate, or with a minute ciliate<br />

fringe; culms to 1.8 m long; cauline blades 4-12 mm wide, usually glabrous; spikelets 1.6-2.8<br />

mm long.<br />

26 Spikelets 2.4-2.8 mm long, long-acuminate, usually < 0.7 mm wide; fertile lemma often<br />

conspicuously stipitate........................................................... P. rigidulum var. elongatum<br />

26 Spikelets 1.6-2.5 mm long, short-acuminate, usually > 0.7 mm wide; fertile lemma<br />

estipitate to short stipitate.<br />

27 Culms to 1 m long; mature panicle ½ to nearly as wide as long, the branches<br />

ascending to spreading; spikelets 1.6-2.2 mm long......... P. rigidulum var. rigidulum<br />

27 Culms to 1.8 m long; mature panicle < 1/3 as wide as long, the branches erect;<br />

spikelets 2.0-2.5 mm long.............................................P. rigidulum var. condensum<br />

Panicum amarum Elliott var. amarulum (A.S. Hitchcock & Chase) P.G. Palmer, Southern Seabeach Grass. Cp (GA, NC,<br />

SC, VA): coastal dunes and shores, sandflats, and sandhills; rare. July-November. NJ s. to FL and West Indies, w. to TX and<br />

Mexico; restricted to the Coastal Plain except for WV. Although well-marked individuals of var. amarulum and var. amarum are<br />

quite distinctive, only the number and structure of first glume nerves appears to be a constant over the range of the two taxa<br />

(Palmer 1975). Primarily a coastal plant, var. amarulum has been found in the Sandhills of NC (Richmond Co.). Blomquist<br />

1948 says this taxon "does not seem to grow naturally in North Carolina." [= K, Z; = P. amarulum A.S. Hitchcock & Chase –<br />

RAB, C, F, G, HC, S; = P. amarum ssp. amarulum (A.S. Hitchcock & Chase) Freckmann & Lelong – FNA; not Panicum]<br />

Panicum amarum Elliott var. amarum, Bitter Seabeach Grass. Cp (GA, NC, SC, VA): coastal dunes and shores; common.<br />

August-November. CT s. to FL, w. to TX; restricted to the coast. See note under var. amarulum. [= K, Z; = P. amarum – RAB,<br />

C, F, G, HC, S; = P. amarum ssp. amarum – FNA; not Panicum]<br />

Panicum anceps Michaux var. anceps, Beaked Panic Grass. Mt, Pd, Cp (NC, SC, VA), {GA}: moist sandy woods,<br />

swamps, sloughs, roadsides, fields, waste places; common. June-October. NJ w. to IL, s. to FL and TX. The sheaths of var.<br />

anceps are glabrous to pilose, while those of var. rhizomatum are often villous; the leaves of var. rhizomatum also tend to be<br />

hairier. [= RAB, F, G, Z; < P. anceps – C, GW, K, W; = P. anceps ssp. anceps – FNA; = P. anceps – HC, S; not Panicum]<br />

Panicum anceps Michaux var. rhizomatum (A.S. Hitchcock & Chase) Fernald, Small Beaked Panic Grass. Cp (GA, NC,<br />

SC, VA): moist to dry sandy or loamy pinelands, ditches; common (VA Watch List). July-October. Se. VA and KY s. to FL<br />

and TX. See note under var. anceps. [= RAB, F, G, Z; < P. anceps – C, GW, K; = P. anceps ssp. rhizomatum (A.S. Hitchcock<br />

& Chase) Freckmann & Lelong – FNA; = P. rhizomatum A.S. Hitchcock & Chase – HC, S; not Panicum]<br />

Panicum capillare Linnaeus, Old-witch Grass, Tumbleweed, Tickle Grass. Mt, Pd (GA, NC, SC, VA), Cp (VA): open<br />

sandy or stony soil, fields, roadsides, waste places, often weedy in cultivated soil; common (rare in SC). August-November. E.<br />

to c. Canada, s. to FL and TX; Bermuda. Plants formerly known as P. capillare var. occidentale Rydberg, ranging from Canada<br />

south to NJ, WV, KY, TX, and CA, are distinguished by long-acuminate spikelets 2.5-4 mm long that are mostly subsessile or<br />

short-pedicelled. In our region, P. capillare has short-acuminate spikelets 1.8-2.8 mm long, mostly on long pedicels. [= RAB,<br />

K, S, Z; < P. capillare – C, Y (also see P. gattingeri); > P. capillare var. capillare – F, HC, W; = P. capillare ssp. capillare –<br />

FNA; = P. capillare var. agreste Gattinger – G; Panicum s.s.]<br />

Panicum dichotomiflorum Michaux var. dichotomiflorum, Spreading Panic Grass, Fall Panic Grass. Mt, Pd, Cp (GA, NC,<br />

SC, VA): marshy shores, exposed wet soils, alluvial deposits in floodplain forests, spoil banks, ditches; common. July-October.<br />

E. Canada w. to SD, s. to FL and TX; also in the Bahamas (Sorrie & LeBlond 1997). P. dichotomiflorum var. puritanorum<br />

ranges along the coast from s. NH to DE, and occurs inland in n. IN. It is distinguished by ovoid to ellipsoid, abruptly short-

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