flora neotropica - CNCFlora

Systematic Treatment 65
Couepia glandulosa Miquel, Stirp. surinam. select. 28.
1851. Type. Suriname (fl), Hostmann & Kappler 859
(holotype, U; isotypes, BM, GH, GOET, K, LE, M,
P, S).
Moquilea glandulosa (Miquel) Walpers, Ann. 2: 463.
1852.
Couepia myrtifolia Bentham ex Hooker f., Fl. bras.
14(2): 44. 1867. Type. Venezuela. Amazonas: San
Carlos (fl), Spruce 3681 (holotype, K; isotypes, BM,
BR, F, GH, GOET, LD, LE, NY, P).
Leaf lamina chartaceous to thinly coriaceous,
5-10.5 cm long, the lower surface glabrous or
sparsely lanate-pubescent; petioles 3-6 mm long.
Inflorescences of terminal and axillary panicles
with small few-flowered branches, in only one or
two axils below the apex of branch. Receptacle
5-10 mm, subcampanulate to narrowly cylindrical.
Plants of flooded river banks.
6-1c. Couepia guianensis Aublet subsp. divaricata
(Huber) Prance, Brittonia 33: 351.
1981. Fig. 104.
Couepia divaricata Huber, Bol. Mus. Paraense Hist.
Nat. 6: 75. 1910. Type. Brazil. Para: Belem, 14 May
1901 (fl), Huber MG 2030 (holotype, MG; isotype,
BM).
Couepia divaricata Huber var. strictiuscula Huber, Bol.
Mus. Paraense Hist. Nat. 6: 76. 1910. Type. Brazil.
Para: Peixe-Boi, Sep 1908 (fl), R. S. Rodrigues MG
8274 (lectotype, MG; isolectotypes, BM, P, RB 15104,
neate at base, apiculate at apex, the acumen 2-
8 mm long, glabrous above, with a short appressed-lanate
caducous pubescence beneath,
midrib prominent on both surfaces, primary veins
12-14 pairs, plane above, prominent beneath,
secondary venation prominulous, more or less
parallel and 90? to primary veins; petioles 4-7
mm long, tomentellous, becoming glabrous with
age, rugulose, canaliculate above, eglandular.
Stipules linear-lanceolate, tomentellous, to 6 mm
long, caducous. Inflorescences of terminal and
axillary little-branched panicles with central
rachis and short 2-3 flowered branches, or of
racemes, the rachis and branches puberulous.
Receptacle cylindrical, 7-8 mm long, sparsely
gray-puberulous externally, glabrous within except
for deflexed hairs at throat; pedicels ca. 1
mm long. Calyx lobes five, rounded, 2.5 mm
long, puberulous externally. Petals five, white,
the margins ciliate. Stamens 20-23, unilateral,
the arc of the circle opposite to them toothed.
Ovary tomentose. Style pubescent on lower portion
only. Fruit ovoid, ca. 5 cm long, 4 cm broad;
exocarp hard and smooth or crustaceous when
dry, glabrous; mesocarp woody, 6-10 mm thick
when dry; endocarp thin and bony, glabrous
within.
Distribution (Fig. 97). Western Amazonia.
U).
Parinari krukovii Gleason, Bull. Torrey Bot. Club 55:
353. 1933. Type. Brazil. Terr. Rondania: Rio Machado,
Tabajara, Nov-Dec 1931 (fl), Krukoff 1362
(holotype, NY; isotypes, K, MICH).
Representative specimens examined. PERU. LORETO:
Prov. Requena, Rio Ucayali, Arboretum Jenaro Herrera,
Jul-Sep 1974 (fr), Bernardi 7-84 (G, NY), 27 Aug
1976 (fl bud), Bernardi 16219 (G, NY).
BRAZIL. AMAZONAS: Mun. Humaita, Livramento,
Oct-Nov 1934 (fr), Krukoff6635 (NY, US); Tres Casas,
Sep-Oct 1934 (fr), Krukoff6381 (NY, US).
Leaf lamina coriaceous, 6-10.5 cm long, the
lower surface densely lanate-pubescent; petioles
3-6 mm long. Inflorescences of terminal and ax-
illary panicles with small few-flowered branches
in only one or two axils below the apex of branch.
Receptacle 5-7 mm long, cylindrical. Plants of
terra firme.
Specimen citations for these three subspecies
were given in Prance (1981).
6-6.1. Couepia bernardii Prance, Brittonia 33:
347. 1981. Type. Peru. Loreto: Prov. Requena,
Rio Ucayali, Arboretum Jenaro Herrera,
4?55'S, 73?45'W, Tree Number 6-6, Jul-Sep
1974 (fl), Bernardi 6-6 (holotype, NY; isotype,
G).
Tree, the young branches puberulous, soon be-
coming glabrous. Leaf lamina elliptic, charta-
ceous, 9-16 x 3.5-6.5 cm, rounded to subcu-
Local names. Brazil: uchirana. Peru: parinari
blanco.
The closely related species Couepia bernardii,
C. obovata, C. reflexa, and C. sandwithii form a
superspecies (see Prance, 1972, for other similar
groups in Couepia). The greater amount of material
available now enables me to group these
species together more clearly than was possible
in 1972. They are characterized by the almost
racemose inflorescence with small groups of
flowers on short branches, thus technically panicles,
the long tubular receptacle that is curved
in bud at the apex, and the reticulate leaf undersurface
with parallel secondary venation in
all species except C. reflexa. I am maintaining
all four species at present since they do appear
to be distinct. Since C. reflexa is only known by