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flora neotropica - CNCFlora

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Systematic Treatment 81<br />

16. Stamens 3; bracteoles coriaceous, ovate; petioles 3-4 mm long.<br />

16. H. davisii.<br />

16. Stamens 4; bracteoles membranous, lanceolate; petioles 1-2 mm long.<br />

17. H. subglanduligera.<br />

2. Leaves hirsute beneath at least on principal venation, usually more generally so.<br />

17. Calyx lobes with small stipitate glands.<br />

18. Leaves orbicular to ovate-elliptic, 3-6.5 cm long, retuse to mucronate at apex (rarely acuminate);<br />

trunk with corky bark; receptacle pubescent within to base. 18. H. ciliata.<br />

18. Leaves oblong to ovate, 4.5-23 cm long, acute to acuminate at apex; trunk with thin bark;<br />

receptacle glabrous within except at throat.<br />

19. Venation of leaves prominulous above; inflorescence subcorymbose, bearing numerous<br />

tightly clustered flowers; stamens 7. 19. H. hoehnei.<br />

19. Venation of leaves impressed or prominulous above; inflorescence lax, spreading, not<br />

subcorymbose, the flowers loosely arranged; stamens 3-5.<br />

20. Leaves subcordate at base, subconduplicate, the lower surface merely sparsely<br />

appressed-pubescent.<br />

29. H. adderleyi.<br />

20. Leaves rounded to cuneate at base, rarely weakly subcordate but never subconduplicate,<br />

the lower surface hirsute. 20. H. glandulosa.<br />

17. Calyx lobes eglandular.<br />

21. Venation of leaves impressed above, or the leaf surface bullate.<br />

22. Inflorescence and flowers rufous-tomentose; bracts bearing very few sessile glands, these<br />

often obscured by pubescence.<br />

42. H. obidensis.<br />

22. Inflorescence and flowers tomentellous to puberulous, not rufous; bracts bearing many<br />

obvious stalked or sessile glands.<br />

23. Leaf surface usually distinctly bullate, the base usually subcordate. 21. H. bullata.<br />

23. Leaf upper surface plane, but with impressed venation, the base rounded to subcuneate.<br />

24. Bracteolar glands ca. 0.5 mm in diam., sessile or slightly stipitate; bracteoles<br />

distinct, ovate. 22. H. americana.<br />

24. Bracteolar glands ca. 0.1 mm in diam., on slender stalks; bracteoles reduced<br />

to a mass of glands with a slightly connate base. 27. H. tentaculata.<br />

21. Venation of leaves prominulous or plane above, the leaf surface never bullate.<br />

25. Stamens 3.<br />

26. Bracteolar glands large, numerous, apparent; inflorescence tomentellous, manyflowered;<br />

exocarp glabrescent; endocarp thin and bony. 22. H. americana.<br />

26. Bracteolar glands small, few; inflorescence hispid, few-flowered; exocarp tomentose;<br />

endocarp thick and fibrous. 23. H. guatemalensis.<br />

25. Stamens 4-7.<br />

27. Exocarp tomentose; endocarp thick and fibrous; bracts with few inconspicuous<br />

stipitate or sessile glands.<br />

28. Inflorescence hispid, the flowers lax; bracts with small short-stipitate glands;<br />

stamens 4. 23. H. guatemalensis.<br />

28. Inflorescence tomentellous, bearing small clusters of flowers on a long thick<br />

rachis; bracts with a few large sessile glands or a single terminal gland; stamens<br />

6. 24. H. eriandra.<br />

27. Exocarp glabrescent; endocarp thin and bony; bracts with many conspicuous stipitate<br />

glands, except in H. liesneri.<br />

29. Inflorescence hispid-hirsute; bracts with short-stalked glands arising from their<br />

margins, the secretory tip small, ellipsoid.<br />

30. Inflorescences predominantly terminal; leaf bases rounded to weakly subcordate.<br />

25. H. paniculata.<br />

30. Inflorescences predominantly axillary; leaf bases deeply cordate.<br />

26. H. deflexa.<br />

29. Inflorescence puberulous to tomentellous; bracts either reduced to numerous,<br />

long-stalked clavate glands arising from the junction of the pedicels with the<br />

stem, or ovate with glands borne on their margins, the secretory tip large,<br />

flattened, forming an expanded head.<br />

31. Glands arising from junction of pedicel and stem, with a long slender<br />

stalk and a small secretory tip, the whole appearing clavate; young stems<br />

puberulous or sparsely tomentellous. 28. H. macrophylla.<br />

31. Glands arising from the bracts, each gland with a thick stalk and a large<br />

secretory tip, flattened to form an expanded head; young stems with short<br />

dense compact puberulence.

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