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Habitat use and population dynamics of the Azure-Winged Magpie ...

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(Dendrocopus spp. <strong>and</strong> Picus canus) were present <strong>and</strong>/ or feeding in <strong>the</strong> same area. C. cyanus<br />

may exert some aggressive dominant behaviour (e.g. chasing <strong>of</strong>f o<strong>the</strong>r species) towards o<strong>the</strong>r<br />

smaller species such as wagtails (Motacilla spp.) if both are competing for food. Hosono (1975)<br />

found similar behaviour in his study in Nagano, Japan.<br />

No obvious territorial behaviours towards conspecifics <strong>and</strong> o<strong>the</strong>r species were observed in this<br />

colony <strong>of</strong> C. cyanus, even during <strong>the</strong> breeding season. I also did not observe any sort <strong>of</strong> obvious<br />

aggression amongst group members. Reports from observations in Iberia <strong>and</strong> Japan show a mix<br />

<strong>of</strong> results, with some <strong>population</strong>s showing territorial behaviours, such as chasing (Komeda et al.,<br />

1987; Snow et al., 1998; S Imanishi, pers. comm.), but o<strong>the</strong>rs not (Valencia et al., 2000). This<br />

non-territorial behaviour may stem from relatedness <strong>of</strong> group members, but kinship has not yet<br />

been studied in <strong>the</strong>se <strong>population</strong>s.<br />

Behavioural observations were limited by <strong>the</strong> difficulty <strong>of</strong> observing C. cyanus when <strong>the</strong>y<br />

foraged in dense bushes (e.g. Padus asiatica <strong>and</strong> Salix spp.). There is no o<strong>the</strong>r way around this<br />

problem, or to improve it without alarming <strong>the</strong> whole colony, apart from conducting many, or<br />

chance observations regarding <strong>the</strong> activities <strong>of</strong> C. cyanus in dense scrub. Observation events <strong>of</strong><br />

areas on <strong>the</strong> true right <strong>of</strong> Eroo River (i.e. <strong>the</strong> Main Breeding Area <strong>and</strong> Sharlan Meadows) were<br />

limited when <strong>the</strong> water level was high, as it was impossible to cross by foot or horse. Therefore,<br />

when this happened (about two to three days in 2009 <strong>and</strong> one week in 2010), observations were<br />

limited to <strong>the</strong> true left <strong>of</strong> <strong>the</strong> Eroo, on <strong>the</strong> true right river bank <strong>and</strong> ca. 20 metres inl<strong>and</strong> from <strong>the</strong><br />

true right <strong>of</strong> <strong>the</strong> river.<br />

4.2 Radio-tracking<br />

VHF radio tracking was employed in this study beca<strong>use</strong> it is by far <strong>the</strong> cheapest tracking method<br />

<strong>and</strong> had reasonable accuracy. As far as I can determine, no VHF radio tracking research has been<br />

published for this species before. Overall, <strong>the</strong> radio-tracking attempts were not successful on C.<br />

cyanus, as I had difficulties getting accurate signal positioning from transmitters <strong>and</strong> securing<br />

transmitters onto <strong>the</strong> tail-fea<strong>the</strong>rs <strong>of</strong> C. cyanus.<br />

Using VHF radio tracking is not without its problems. Tests <strong>of</strong> radiotelemetry accuracy have<br />

shown signal reception <strong>and</strong> location accuracy to be affected by several factors, amongst <strong>the</strong>m<br />

reflected signals (“signal bounce”) <strong>and</strong> attenuation resulting from vegetation cover (Wi<strong>the</strong>y et al.,<br />

2001), which I think were <strong>the</strong> main reasons for difficulties in getting signals from <strong>the</strong> radio-<br />

61

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