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Fauna of New Zealand 69 - Landcare Research

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30<br />

11(10) Head, pronotum, and dark areas <strong>of</strong> elytra blackish<br />

brown. Elytra (Fig. 158): pale markings narrow;<br />

humeral lunule separated from marginal line; middle<br />

band jagged. [Body length 8.4–10.7 mm; occurs on<br />

black ironsand; northern North Island (western AK<br />

and WO).] .......................(p. 34)... campbelli (Broun)<br />

—Head, pronotum, and dark areas <strong>of</strong> elytra greenish<br />

bronze. Elytra (Fig. 159): pale markings very wide;<br />

humeral lunule fused with marginal line; middle<br />

band not jagged. [Body length 8.5–10.3 mm; occurs<br />

on yellow sand; northern North Island (western ND<br />

and AK).] .....................(p. 35)... perhispida (Broun)<br />

Zecicindela hamiltoni (Broun, 1921) E new<br />

combination<br />

Figures 51, 148; Map p. 184<br />

Common name. Hamilton’s Tiger Beetle.<br />

Cicindela (Neocicindela) hamiltoni: Larochelle & Larivière,<br />

2001: 37 and 2007a: 110.<br />

Neocicindela hamiltoni: Lorenz, 2005: 61; Brzoska, 2006: 29;<br />

Cassola & Moravec, 2010: 12.<br />

Description. Body length: 8.5–9.2 mm. Head and pronotum<br />

greenish; dark areas <strong>of</strong> elytra dark brown. Pale<br />

markings <strong>of</strong> elytra cream-coloured, narrow, immaculate;<br />

humeral lunule incomplete, fused with marginal line; middle<br />

band incomplete, almost absent, not extending beyond<br />

base <strong>of</strong> apical lunule; marginal line fused with apical<br />

lunule. Antennae and legs entirely dark. Elytral sculpture:<br />

dark areas granulate, densely covered with minute green<br />

punctures, with small green foveae in subsutural row and<br />

humeral area. Dark areas <strong>of</strong> elytra rather dull, with slight<br />

greenish metallic lustre. Head. Labrum quadrisetose anteriorly.<br />

Antennal segment 1 (scape) with a single setiferous<br />

pore. Thorax. Pronotum (Fig. 148) subcordate; sides<br />

moderately rounded; lateral margins double. Proepisterna<br />

and metepisterna setose. Legs. Tarsal claws much shorter<br />

than tarsal segment 5 (about half its length). Elytra. Sides<br />

widening behind middle. Apices not serrulate. Abdomen.<br />

Sterna glabrous. Aedeagus (Fig. 51).<br />

References. Larochelle & Larivière, 2001: 37 (as Cicindela<br />

(Neocicindela) hamiltoni; catalogue; biology, dispersal<br />

power, ecology, geographic distribution, references);<br />

Brzoska, 2006: 29 (as Neocicindela hamiltoni; ecology);<br />

Cassola & Moravec, 2010: 12 (as Neocicindela hamiltoni;<br />

ecology, geographic distribution).<br />

Zecicindela helmsi novaseelandica (Horn, 1892) E<br />

new combination and status<br />

Figure 149; Map p. 184<br />

Common name. <strong>New</strong> <strong>Zealand</strong> Tiger Beetle.<br />

Larochelle & Larivière (2013): Carabidae (Insecta: Coleoptera)<br />

Cicindela novaseelandica: as a junior synonym <strong>of</strong> Cicindela<br />

(Neocicindela) helmsi (Larochelle & Larivière 2001: 37).<br />

Neocicindela novaseelandica: resurrected from synonymy with<br />

Neocicindela helmsi by Cassola & Moravec, 2010: 13.<br />

Description. Body length: 9.2–11.3 mm. Head and<br />

pronotum aeneous or bronze, with greenish tinge; dark<br />

areas <strong>of</strong> elytra purplish-brown, with greenish tinge. Pale<br />

markings cream-coloured, moderately wide; apical lunule<br />

maculate; humeral lunule complete, usually widely separated<br />

from marginal line; middle band complete, angular,<br />

not extending beyond base <strong>of</strong> apical lunule; marginal line<br />

usually widely separated from apical lunule. Antennae and<br />

legs entirely dark. Elytral sculpture: dark areas granulate,<br />

densely covered with minute blue punctures ringed with<br />

golden-green, with sparse large bluish foveae with golden<br />

centre (foveae deep giving uneven elytral surface). Dark<br />

areas <strong>of</strong> elytra dull, with slight greenish, aeneous or bronze<br />

metallic lustre. Head. Labrum quadrisetose anteriorly.<br />

Antennal segment 1 (scape) with 9–16 setiferous pores.<br />

Thorax. Pronotum (Fig. 149) subcordate; sides moderately<br />

rounded; lateral margins double. Proepisterna and metepisterna<br />

setose. Legs. Tarsal claws much shorter than tarsal<br />

segment 5 (about half its length). Elytra. Sides strongly<br />

widening behind middle. Apices not serrulate. Abdomen.<br />

Sterna setose. Aedeagus. As in nominal subspecies.<br />

References. Larochelle & Larivière, 2001: 37 (as Cicindela<br />

(Neocicindela) helmsi; catalogue; biology, dispersal<br />

power, ecology, geographic distribution, references);<br />

Cassola & Moravec, 2010: 13–16 (as Neocicindela novaseelandica;<br />

ecology, geographic distribution, taxonomy).<br />

Remarks. Zecicindela helmsi novaseelandica is closest<br />

to Zecicindela helmsi halli, from which it can be<br />

distinguished by a combination <strong>of</strong> morphological and<br />

eco-geographic characters: elytra with humeral lunule<br />

usually widely separated from marginal line and apical<br />

lunule usually widely separated from marginal line;<br />

occurring away from water, on roads and trails <strong>of</strong> the<br />

Central Volcanic Plateau (TO) <strong>of</strong> the North Island. Other<br />

subspecies <strong>of</strong> Z. helmsi are associated with South Island<br />

riverbanks. Cassola & Moravec (2010) provided photos<br />

<strong>of</strong> the left elytron <strong>of</strong> the lectotype and paralectotype <strong>of</strong><br />

Cicindela novaseelandica.<br />

See also Remarks under Z. helmsi helmsi.<br />

Zecicindela helmsi halli (Broun, 1917) E new<br />

combination and status<br />

Figure 150; Map p. 184<br />

Common name. Hall’s Tiger Beetle.<br />

Cicindela halli: as a junior synonym <strong>of</strong> Cicindela (Neocicindela)<br />

helmsi (Larochelle & Larivière, 2001: 37).<br />

Cicindela halli: resurrected from synonymy with Neocicindela

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