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Fauna of New Zealand 69 - Landcare Research

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<strong>Fauna</strong> <strong>of</strong> <strong>New</strong> <strong>Zealand</strong> <strong>69</strong> 61<br />

anteriorly, moderately sinuate posteriorly; lateral beads<br />

narrow; lateral depressions absent; each anterolateral setiferous<br />

puncture situated just in front <strong>of</strong> middle and touching<br />

lateral bead; posterolateral angles usually rectangular,<br />

sometimes acute and projecting laterally, not denticulate;<br />

laterobasal foveae well defined, finely punctate, shallow,<br />

narrow, oblong; base slightly arcuate, about as wide as<br />

apex, moderately narrower than elytral base. Prosternum<br />

impunctate, unwrinkled. Proepisterna impunctate or barely<br />

punctate, unwrinkled. Metepisterna impunctate or barely<br />

punctate. Elytra. Widest in front <strong>of</strong> middle. Shoulders<br />

moderately rounded, denticulate. Sides strongly rounded.<br />

Scutellar striole impunctate. Striae mostly complete (stria<br />

7 obsolete in basal half; complete in other species), shallow,<br />

finely punctate (coarsely punctate in other species).<br />

Intervals depressed, becoming slightly convex apically.<br />

Sutural apices angular. Abdomen. Sterna impunctate<br />

(punctate in other species). Last visible sternum (sternum<br />

VII): male with four apical ambulatory setae; female with<br />

four apical and two medial ambulatory setae. Aedeagus.<br />

Lateral view (Fig. 76): strongly arcuate; apex moderately<br />

wide, moderately rounded, not projecting dorsoventrally,<br />

with a posterodorsal notch. Dorsal view: asymmetrical<br />

(ostium deflected to the right); apex deflected to the right.<br />

Material examined. 71 specimens (AMNZ, BMNH,<br />

CMNZ, JNNZ, LUNZ, NZAC).<br />

Geographic distribution (Map p. 181). South Island:<br />

CO, DN, OL, SL.<br />

Ecology. Lowland, montane, subalpine, alpine. Wet forests<br />

(beech, broadleaf, podocarp), tussock grasslands, and<br />

alpine meadows. Gregarious. Shaded (mostly) or open<br />

ground; wet soil. Nocturnal; active at night on trees; hides<br />

during the day under (mostly) and in logs, also in leaf litter.<br />

Biology. Seasonality: September to March, June, August.<br />

Tenerals: January-February. Predacious (based on mouthpart<br />

morphology).<br />

Dispersal power. Elytra fused basally along suture. Subapterous.<br />

Moderate runner. Regular climber (on trees).<br />

Vagility limited by flight incapacity.<br />

Reference. Larochelle & Larivière, 2001: 68 (biology,<br />

dispersal power, ecology, geographic distribution).<br />

Remark. Examination <strong>of</strong> the holotype <strong>of</strong> Tarastethus<br />

fovealis revealed it to be conspecific with Tarastethus<br />

southlandicus.<br />

Tarastethus sirvidi E new species<br />

Figures 77, 122, 217; Map p. 181<br />

Tarastethus sirvidi Larochelle & Larivière, new species. Holotype:<br />

male (NZAC) labelled “NEW ZEALAND SL<br />

Longwood SF [=State Forest], Bald Hill 700m 26.I.1999<br />

Larivière, Larochelle (typed) / Wet subalpine cloud for.<br />

[=forest] (beech): mossy floor. Under beech branches, in the<br />

open (typed) / HOLOTYPE [male symbol] Tarastethus sirvidi<br />

Larochelle & Larivière, 2013 (red label; typed).” Paratypes:<br />

one male and one female (NZAC) from Longwood<br />

Range, bearing blue paratype labels.<br />

Description. Body: length 4.3–5.6 mm. Head, pronotum,<br />

elytra, and abdomen black; margins and suture <strong>of</strong> elytra<br />

reddish; antennae, palpi, and tarsi pale yellowish; tibiae<br />

and femora infuscate. Head. Finely and sparsely punctate<br />

in and between frontal furrows, unwrinkled dorsally,<br />

slightly narrower across eyes than pronotal apex. Labrum<br />

moderately transverse, subtruncate anteriorly. Antennae:<br />

segment 1 (scape) moderately long, about 1.5× longer<br />

than its maximum width. Frontal furrows wide, shallow.<br />

Eyes smaller than in other species, moderately convex;<br />

two supraorbital setiferous punctures on inner side <strong>of</strong><br />

each eye. Interocular fovea absent or almost so. Tempora<br />

convex, moderately long (about half as long as eyes).<br />

Mentum: medial tooth slightly shorter than lateral lobes.<br />

Paraglossae membranous, prominent, much longer than<br />

ligula, slightly converging anteriorly. Thorax. Pronotum<br />

(Fig. 122) strongly convex, coarsely punctate across base,<br />

finely punctate along lateral beads, impunctate on disc,<br />

unwrinkled, strongly cordate; apex straight; anterolateral<br />

angles poorly developed, obtusely rounded; sides strongly<br />

rounded anteriorly, moderately sinuate posteriorly; lateral<br />

beads slightly widening from apex to base; lateral depressions<br />

absent; each anterolateral setiferous puncture<br />

situated just in front <strong>of</strong> middle and touching lateral bead;<br />

posterolateral angles usually rectangular, sometimes acute<br />

and projecting laterally, not denticulate; laterobasal foveae<br />

well defined, finely punctate, deep, wide, oblong, elongate<br />

anteriorly; base more or less straight, about as wide as<br />

apex, moderately narrower than elytral base. Prosternum<br />

punctate, unwrinkled. Proepisterna punctate, unwrinkled.<br />

Metepisterna punctate. Elytra. Widest in front <strong>of</strong> middle.<br />

Sides strongly rounded. Shoulders moderately rounded,<br />

denticulate. Scutellar striole absent. Striae complete,<br />

moderately deep, coarsely punctate. Intervals slightly<br />

convex throughout. Sutural apices angular. Abdomen.<br />

Sterna coarsely and sparsely punctate. Last visible sternum<br />

(sternum VII): male with four apical and two subapical<br />

ambulatory setae; female with four apical and four medial<br />

ambulatory setae. Aedeagus. Lateral view (Fig. 77):<br />

strongly arcuate; apex very wide, truncate, not projecting<br />

dorsoventrally, notched dorsally only. Dorsal view: symmetrical<br />

(ostium expanding equally towards sides); apex<br />

deflected to the left.

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