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Fauna of New Zealand 69 - Landcare Research

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70<br />

Biology. Seasonality: October to June, August. Tenerals:<br />

October, January, March. Predacious (based on mouthpart<br />

morphology).<br />

Dispersal power. Elytra fused basally along the suture.<br />

Subapterous. Moderate runner. Vagility limited by flight<br />

incapacity.<br />

Reference. Larochelle & Larivière, 2001: 67 (biology,<br />

dispersal power, ecology, geographic distribution).<br />

Remark. Examination <strong>of</strong> the holotype <strong>of</strong> Tarastethus cinctus<br />

revealed it to be conspecific with Trichopsida oxygona.<br />

Trichopsida optata (Broun, 1917) E new combination<br />

Figures 129, 224; Map p. 182<br />

Tarastethus optatus Broun, 1917: 3<strong>69</strong>. Holotype: female<br />

(BMNH) labelled “Type (circular red-bordered label;<br />

typed) / 3815- (hand-written) / <strong>New</strong> <strong>Zealand</strong>. Broun Coll.<br />

Brit. Mus. 1922-482. (white label with red horizontal line;<br />

typed) / Mount Dick. 17-3-1914. (hand-written) / Tarastethus<br />

optatus-[female symbol]- (hand-written).” Condition:<br />

Good (terminal segment <strong>of</strong> right antenna missing).<br />

Molopsida optata: Britton, 1940: 277.<br />

Description. Body: length 5.6–5.9 mm. Head, pronotum,<br />

elytra brownish black; sides <strong>of</strong> pronotum dark or somewhat<br />

paler anterolaterally; sides <strong>of</strong> elytra narrowly pale<br />

yellowish; abdomen dark brownish; antennae, palpi, and<br />

femora pale red; tibiae and tarsi infuscate. Microsculpture<br />

very transverse (with microlines), feeble on head<br />

and pronotum, strong on elytra. Iridescence absent on<br />

head and pronotum, strong on elytra. Very shiny. Head.<br />

Impunctate and unwrinkled dorsally (as in T. oxygona),<br />

much narrower across eyes than pronotal apex. Mandibles<br />

short. Labrum moderately transverse and emarginate<br />

anteriorly. Antennae: segment 1 (scape) moderately long,<br />

about 1.5× longer than its maximum width; segments 1–3<br />

glabrous (excluding apical setae). Frontal furrows wide,<br />

rather deep. Eyes normally developed, moderately large,<br />

slightly convex. Tempora convex, moderately long (about<br />

half as long as eyes). Mentum: medial tooth acute apically,<br />

slightly shorter than lateral lobes. Ligula narrow apically.<br />

Paraglossae membranous, prominent, much longer than<br />

ligula. Palpi: terminal segment fusiform, not elliptical,<br />

sparsely setulose; penultimate maxillary segment apically<br />

setulose. Thorax. Pronotum (Fig. 129) moderately convex,<br />

finely punctate across base, feebly wrinkled along median<br />

longitudinal impression, strongly transverse, strongly<br />

cordate; apex rather straight; anterolateral angles poorly<br />

developed, obtusely rounded; sides strongly rounded<br />

anteriorly, oblique and slightly sinuate posteriorly; lateral<br />

beads narrow; lateral depressions absent; a single setiferous<br />

puncture on each side (anteriorly), situated just in front<br />

<strong>of</strong> middle and touching lateral bead; posterolateral angles<br />

Larochelle & Larivière (2013): Carabidae (Insecta: Coleoptera)<br />

obtuse, denticulate; laterobasal foveae well defined, finely<br />

punctate, shallow, narrow, elongate medially; posterior<br />

bead complete; base emarginate medially, slightly oblique<br />

laterally, slightly wider than apex, moderately narrower<br />

than elytral base. Prosternum impunctate, unwrinkled.<br />

Proepisterna impunctate, unwrinkled. Metepisterna impunctate.<br />

Elytra. Moderately convex, subovate, widest<br />

about middle. Basal margin complete. Shoulders obtuse,<br />

not denticulate. Sides moderated rounded. Scutellar setiferous<br />

pore inserted at junction <strong>of</strong> striae 1+2. Scutellar striole<br />

barely visible, almost impunctate. Striae mostly complete<br />

(striae 6–7 obsolete basally), shallow, finely punctate; stria<br />

3 with two small setiferous punctures. Intervals depressed.<br />

Apices angular-rounded. Abdomen. Sterna impunctate,<br />

unwrinkled. Last visible sternum (sternum VII): male<br />

unknown; female with four to six apical and three to five<br />

medial ambulatory setae. Aedeagus and Parameres.<br />

Male unknown.<br />

Material examined. Six specimens (BMNH, NZAC).<br />

Geographic distribution (Map p. 182). South Island:<br />

OL─Mount Dick. North Mavora Lake. South Mavora<br />

Lake.<br />

Ecology. Lowland (hills), subalpine. Wet forests (beech).<br />

Shaded ground; mossy soil. Probably nocturnal and hiding<br />

during the day in moss. The species is possibly epigean<br />

(living on the surface <strong>of</strong> the ground). Four specimens were<br />

collected by pit trapping.<br />

Biology. Seasonality: January, March. Predacious (based<br />

on mouthpart morphology).<br />

Dispersal power. Elytra fused in basal half. Subapterous.<br />

Moderate runner. Vagility limited by flight incapacity.<br />

Trichopsida erwini E new species<br />

Figures 84, 130, 225; Map p. 181<br />

Trichopsida erwini Larochelle & Larivière, new species. Holotype:<br />

male (NZAC) labelled “Mt Domett NN 1463m 1 Dec<br />

71 J.S. Dugdale (typed) / mat plants (typed) / HOLOTYPE<br />

[male symbol] Trichopsida erwini Larochelle & Larivière,<br />

2013 (red label; typed).” Paratypes: one male (NZAC),<br />

three females (LUNZ, MONZ, NZAC) from the same locality<br />

as the holotype, bearing blue paratype labels.<br />

Description. Body: length 3.2–5.2 mm. Head reddish;<br />

pronotum dark brown medially; elytra dark brown, with<br />

sides and apex brownish red; abdomen mostly yellowish,<br />

brownish black medially; antennae, palpi, and legs entirely<br />

pale yellowish. Microsculpture absent. Iridescence absent.<br />

Very shiny. Head. Impunctate and sparsely wrinkled<br />

dorsally, much narrower across eyes than pronotal apex.<br />

Mandibles moderately long. Labrum slightly transverse,<br />

truncate or slightly emarginate anteriorly. Antennae: segment<br />

1 (scape) moderately long, about 1.5× longer than its

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