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Fauna of New Zealand 69 - Landcare Research

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<strong>Fauna</strong> <strong>of</strong> <strong>New</strong> <strong>Zealand</strong> <strong>69</strong> 71<br />

maximum width; segments 1–3 glabrous (excluding apical<br />

setae). Frontal furrows narrow, shallow. Eyes reduced,<br />

very small, slightly convex. Tempora convex, moderately<br />

long (about half as long as eyes). Mentum: medial tooth<br />

acute apically, slightly shorter than lateral lobes. Ligula<br />

narrow apically. Paraglossae membranous, prominent,<br />

much longer than ligula. Palpi: terminal segment fusiform,<br />

not elliptical, sparsely setulose; penultimate maxillary<br />

segment apically setulose. Thorax. Pronotum (Fig.<br />

130) moderately convex, coarsely and sparsely punctate<br />

across base, unwrinkled, moderately transverse, strongly<br />

cordate; apex strongly arcuate laterally, straight medially;<br />

anterolateral angles strongly developed, obtuse or obtusely<br />

rounded; sides strongly rounded anteriorly, strongly sinuate<br />

posteriorly; lateral beads strongly widening from apex<br />

to base; lateral depressions widening posteriorly; a single<br />

setiferous puncture on each side (anteriorly), situated well<br />

in front <strong>of</strong> middle and touching lateral bead; posterolateral<br />

angles subrectangular or acute and projecting laterally,<br />

not denticulate; laterobasal foveae ill-defined, coarsely<br />

punctate, deep, wide, elongate medially; posterior bead<br />

complete; base rather straight, slightly narrower than apex,<br />

much narrower than elytral base. Prosternum impunctate,<br />

unwrinkled. Proepisterna punctate, unwrinkled. Metepisterna<br />

impunctate. Elytra. Strongly convex, ovate, widest<br />

about middle. Basal margin complete. Shoulders rounded,<br />

denticulate. Sides strongly rounded. Scutellar setiferous<br />

pore inserted at junction <strong>of</strong> striae 1+2. Scutellar striole<br />

absent. Striae complete, striae 1–5 moderately deep, striae<br />

6–7 shallower, with moderately coarse punctation; stria 3<br />

with two setiferous punctures. Intervals slightly convex.<br />

Sutural apices angular. Abdomen. Sterna impunctate,<br />

unwrinkled. Last visible sternum (sternum VII): male with<br />

two apical ambulatory setae; female with four apical and<br />

two or four medial ambulatory setae. Aedeagus. Lateral<br />

view (Fig. 84): strongly arcuate; apex moderately wide,<br />

subtriangular, not projecting dorsoventrally. Dorsal view:<br />

asymmetrical (ostium deflected to the right); apex deflected<br />

to the right. Parameres. Glabrous.<br />

Material examined. 44 specimens (LUNZ, MONZ,<br />

NZAC).<br />

Geographic distribution (Map p. 181). South Island:<br />

BR─Lake Rotoroa. NN─Canaan. Cobb Reservoir. Cobb<br />

Valley. Karamea (9 miles north). Mount Arthur (Flora Hut,<br />

Flora Stream, Flora Track, Tableland). Mount Domett.<br />

Oparara River.<br />

Ecology. Lowland, montane, subalpine, alpine. Wet forests<br />

(beech), shrublands, and fellfields. Shaded or open ground;<br />

wet soil. Nocturnal; hides during the day in leaf litter, moss,<br />

and plant mats. Gregarious.<br />

Biology. Seasonality: November to February, April, June.<br />

Tenerals: November. Predacious (based on mouthpart<br />

morphology).<br />

Dispersal power. Elytra fused basally along suture.<br />

Subapterous. Moderate runner. Vagility limited by flight<br />

incapacity.<br />

Remarks. This species is named after our friend and colleague<br />

Terry L. Erwin (Smithsonian Museum, Washington<br />

D.C.) for his authoritative contribution to the study <strong>of</strong><br />

world Carabidae.<br />

Trichopsida erwini is morphologically close to T.<br />

pretiosa and T. paturauensis. In addition to diagnostic<br />

characters <strong>of</strong> the male genitalia, especially the subtriangular<br />

apex <strong>of</strong> the aedeagus, T. erwini has the following<br />

distinguishing features: pronotum strongly cordate, with<br />

sides strongly sinuate and posterolateral angles subrectangular<br />

or acute and projecting laterally; elytral striae<br />

with moderately coarse punctation.<br />

Trichopsida pretiosa (Broun, 1910) E new<br />

combination<br />

Figures 85, 131, 226; Map p. 183<br />

Tarastethus pretiosus Broun, 1910: 6. Holotype: male (BMNH)<br />

labelled “Type (circular red-bordered label; typed) / 3022<br />

(hand-written) / <strong>New</strong> <strong>Zealand</strong>. Broun Coll. Brit. Mus.<br />

1922–482. (white label with red horizontal line; typed) /<br />

Raurimu. Jany. 1909. (hand-written) / Tarastethus pretiosus.<br />

(hand-written).” Condition: Excellent. Paratype: male<br />

(BMNH) labelled “Paratype (circular yellow-bordered label;<br />

typed) / 3022 (hand-written) / <strong>New</strong> Zeal. Broun Coll.<br />

Brit. Mus. 1922–482. (white label with red horizontal line;<br />

typed) / Raurimu. Jany. 1909. (hand-written) / Tarastethus<br />

pretiosus. (hand-written).” Condition: Fair.<br />

Molopsida pretiosa: Britton, 1940: 477.<br />

Description. Body: length 3.9–4.4 mm. Head and pronotum<br />

reddish; pronotum dark brown medially; elytra dark<br />

brown with sides and apex brownish red; abdomen mostly<br />

yellowish, blackish medially; antennae, palpi, and legs<br />

entirely pale yellowish. Microsculpture absent. Iridescence<br />

absent. Very shiny. Head. Impunctate and unwrinkled<br />

dorsally, much narrower across eyes than pronotal apex.<br />

Mandibles moderately long. Labrum moderately transverse,<br />

truncate anteriorly. Antennae: segment 1 (scape)<br />

moderately long, about 1.5× longer than its maximum<br />

width; segments 1–3 glabrous (excluding apical setae).<br />

Frontal furrows wide, deep. Eyes reduced, very small,<br />

slightly convex. Tempora convex, moderately long (about<br />

half as long as eyes). Mentum: medial tooth acute apically,<br />

slightly shorter than lateral lobes. Ligula narrow apically.<br />

Paraglossae membranous, prominent, much longer than<br />

ligula. Palpi: terminal segment fusiform, not elliptical,<br />

sparsely setulose; penultimate maxillary segment apically<br />

setulose. Thorax. Pronotum (Fig. 131) moderately convex,

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