Fauna of New Zealand 69 - Landcare Research

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Neocicindela helmsi: Lorenz, 2005: 61; Brzoska, 2006: 29; Cassola
& Moravec, 2010: 12.
Description. Body length: 8.7–12.5 mm. Head and pronotum
aeneous or bronze, with greenish tinge; dark areas of
elytra purplish-brown, with greenish tinge. Pale markings
of elytra cream-coloured, moderately wide; apical lunule
maculate; humeral lunule complete, narrowly separated
from marginal line; middle band complete, oblique, not
extending beyond base of apical lunule; apical lunule
narrowly separated from marginal line. Antennae and
legs entirely dark. Elytral sculpture: dark areas granulate,
densely covered with minute blue punctures ringed with
golden-green, with sparse small bluish foveae with golden
centre (foveae shallow giving even elytral surface). Dark
areas of elytra dull, with slight greenish, aeneous or bronze
metallic lustre. Head. Labrum quadrisetose anteriorly.
Antennal segment 1 (scape) with 12–16 setiferous pores.
Thorax. Pronotum (Fig. 152) subcordate; sides moderately
rounded; lateral margins double. Proepisterna and
metepisterna setose. Legs. Tarsal claws much shorter
than tarsal segment 5 (about half its length). Elytra. Sides
subparallel. Apices not serrulate. Abdomen. Sterna setose.
Aedeagus (Fig. 52).
References. Larochelle & Larivière, 2001: 37 (as Cicindela
(Neocicindela) helmsi; catalogue; biology, dispersal
power, ecology, geographic distribution, references);
Brzoska, 2006: 29–32 (as Neocicindela helmsi; ecology,
geographic distribution); Cassola & Moravec, 2010: 12–13
(as Neocicindela helmsi; ecology, geographic distribution,
taxonomy).
Remarks. Cassola & Moravec (2010) resurrected Cicindela
novaeseelandica Horn, 1892, from synonymy with
Neocicindela helmsi and provided the new combination
Neocicindela novaeseelandica. These authors also placed
Cicindela halli Broun, 1917, previously a synonym of N.
helmsi, as a junior synonym of N. novaeseelandica. These
taxonomic changes were based on two elytral characters,
the degree of fusion of the pale marginal band and the
number of foveae, and on extremely limited material
representing few populations. No reference was made to
the configuration of the male genitalia but a photo of the
left elytron of the holotype was provided.
Our examination of the large amount of material
identified as N. helmsi in New Zealand collections revealed
a single species with an identical male aedeagus
across four morphological subspecies (Z. helmsi helmsi,
Z. helmsi circumpictoides, Z. helmsi halli, Z. helmsi novaseelandica),
and with geographic distributions that
do not overlap. Results also showed that the diagnostic
characters used by Cassola & Moravec (2010) are highly
variable within and between populations of Z. helmsi.
Zecicindela helmsi helmsi is morphologically close
to Z. helmsi circumpictoides, from which it can be dis-
Larochelle & Larivière (2013): Carabidae (Insecta: Coleoptera)
tinguished by the following morphological characters:
elytra with humeral lunule and apical lunule narrowly
separated from marginal line, and dark areas with sparse
larger shallow foveae.
Zecicindela helmsi helmsi occurs on riverbanks of
the northwestern South Island (NN, BR).
See also Remarks under other subspecies.
Zecicindela feredayi (Bates, 1867) E new combination
Figures 53, 153; Map p. 183
Common name. Fereday’s Tiger Beetle.
Cicindela (Neocicindela) feredayi: Larochelle & Larivière,
2001: 37 and 2007a: 110.
Neocicindela feredayi: Lorenz, 2005: 61; Brzoska, 2006: 32;
Cassola & Moravec, 2010: 17.
Description. Body length: 7.5–9.0 mm. Head, pronotum,
and dark areas of elytra greenish black. Pale markings
of elytra white, narrow, immaculate; humeral lunule
complete, separated or not from marginal line; middle
band complete, angular, extending close to apical lunule;
marginal line fused with apical lunule. Antennae and
legs entirely dark. Elytral sculpture: dark areas granulate,
densely covered with minute blue punctures with
green centre, with sparse small-sized greenish foveae in
humeral area; subsutural row absent. Dark areas of elytra
dull, with slight greenish metallic lustre. Head. Labrum
quadrisetose anteriorly. Antennal segment 1 (scape) with
9–19 setiferous pores. Thorax. Pronotum (Fig. 153) subcordate;
sides rather moderately rounded; lateral margins
double. Proepisterna and metepisterna setose. Legs. Tarsal
claws much shorter than tarsal segment 5 (about half its
length). Elytra. Sides widening behind middle. Apices not
serrulate. Abdomen. Sterna setose. Aedeagus (Fig. 53).
References. Larochelle & Larivière, 2001: 37 (as Cicindela
(Neocicindela) feredayi; catalogue; biology, dispersal
power, ecology, geographic distribution, references);
Brzoska, 2006: 32–33 (as Neocicindela feredayi; ecology,
geographic distribution); Cassola & Moravec, 2010: 17 (as
Neocicindela feredayi; ecology, geographic distribution).
Zecicindela austromontana (Bates, 1878) E new
combination
Figures 54, 154; Map p. 183
Common name. Southern Mountains Tiger Beetle.
Cicindela (Neocicindela) austromontana: Larochelle &
Larivière, 2001: 36 and 2007a: 110.
Neocicindela austromontana: Lorenz, 2005: 61; Brzoska, 2006:
32; Cassola & Moravec, 2010: 17.
Description. Body length: 10.0–12.5 mm. Head, pronotum,
and dark areas of elytra olivaceous black. Pale markings
of elytra white, narrow, immaculate; humeral lunule
incomplete, separated or not from marginal line; middle