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Fauna of New Zealand 69 - Landcare Research

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<strong>Fauna</strong> <strong>of</strong> <strong>New</strong> <strong>Zealand</strong> <strong>69</strong> 77<br />

Sides almost straight. Scutellar setiferous pore inserted at<br />

junction <strong>of</strong> striae 1+2. Scutellar striole absent or present<br />

(uni- or bipunctate). Striae mostly complete (stria 7 obsolete<br />

basally), moderately deep, with moderately coarse<br />

punctation; stria 3 without setiferous punctures. Intervals<br />

depressed in basal half, slightly convex in apical half.<br />

Sutural apices angular. Abdomen. Sterna impunctate,<br />

wrinkled laterally. Last visible sternum (sternum VII):<br />

male with four apical ambulatory setae; female with four<br />

apical and two medial ambulatory setae. Aedeagus. Lateral<br />

view (Fig. 89): moderately arcuate; apex narrow, strongly<br />

projecting dorsally. Dorsal view: asymmetrical (ostium<br />

deflected to the right); apex straight. Parameres. Setulose<br />

(with a few short setae at apex).<br />

Material examined. Nine specimens (JNNZ, NZAC).<br />

Geographic distribution (Map p. 182). South Island:<br />

WN–Keith George Scenic Reserve. Kaitoke. Wellington<br />

(Tinakori Hill, Wilton’s Bush).<br />

Ecology. Lowland (hills). Forests (broadleaf, tree-fern,<br />

pine). Shaded ground. Nocturnal; hides during the day under<br />

deeply embedded stones; also found under a log, with<br />

Mesoponera castanea ants (J. Nunn, personal communication).<br />

The body shape (poorly-developed eyes, inflated<br />

tempora, depressed elytra) and pale colour suggest that the<br />

species is endogean (living in soil crevices or fissures, in<br />

deep humus <strong>of</strong> leaf litter, under well-embedded stones).<br />

Biology. Seasonality: September to January, July–August.<br />

Predacious (based on mouthpart morphology).<br />

Dispersal power. Elytra fused basally along suture.<br />

Subapterous. Moderate runner. Vagility limited by flight<br />

incapacity.<br />

Remarks. This species is named after our friend the<br />

coleopterist John Nunn (Dunedin) who discovered this<br />

species.<br />

Trichopsida nunni is morphologically close to T.<br />

goethei. In addition to diagnostic characters <strong>of</strong> the male<br />

genitalia T. nunni has the following distinguishing features:<br />

microsculpture absent on head, pronotum, and<br />

elytra; tempora very long (about as long as eyes); pronotum<br />

slightly transverse, subrectangular, barely punctate,<br />

unwrinkled, with sides slightly rounded anteriorly, not<br />

sinuate posteriorly, without laterobasal foveae, and with<br />

posterior bead interrupted medially; elytra elongate and<br />

subparallel, with striae distinctly punctate.<br />

Trichopsida debilis (Sharp, 1886) E new combination<br />

Figures 90, 138, 233; Map p. 181<br />

Tarastethus debilis Sharp, 1886: 373. Holotype: male (BMNH)<br />

labelled “Tarastethus debilis. Type D.S. Bealey. N.Z.<br />

Helms. (hand-written on card mount) / Type H.T. (circular<br />

red-bordered label; typed) / Bealey, <strong>New</strong> <strong>Zealand</strong>. Helms.<br />

(typed ) / Sharp Coll. 1905-313. (typed)” Condition: Excellent.<br />

Tarastethus longulus Broun, 1917: 368. Holotype: male<br />

(BMNH) labelled “Type (circular red-bordered label;<br />

typed) / 3813- (hand-written) / <strong>New</strong> <strong>Zealand</strong>. Broun Coll.<br />

Brit. Mus. 1922-482. (white label with red horizontal line;<br />

typed) / Clippings. 28-1-1914. (hand-written) / Tarastethus<br />

longulus (hand-written).” Condition: Excellent. <strong>New</strong> synonym.<br />

Molopsida debilis: Britton, 1940: 477.<br />

Molopsida longula: Britton, 1940: 477.<br />

Description. Body: length 4.2–5.5 mm. Head, pronotum,<br />

and elytra dark reddish; base <strong>of</strong> pronotum and base, suture,<br />

margins, and apex <strong>of</strong> elytra pale yellowish red; abdomen<br />

dark testaceous; antennae, palpi, and femora pale red;<br />

tibiae and tarsi infuscate. Microsculpture moderately transverse,<br />

feeble on head and pronotum, strong on elytra. Iridescence<br />

absent. Very shiny. Head. Coarsely and entirely<br />

punctate, wrinkled dorsally in apical half (unwrinkled in<br />

other species), slightly narrower across eyes than pronotal<br />

apex. Mandibles moderately long. Labrum moderately<br />

transverse and emarginate anteriorly. Antennae: segment<br />

1 (scape) moderately long, about 1.5× longer than its<br />

maximum width; segments 1–3 glabrous (excluding apical<br />

setae). Frontal furrows wide, rather deep. Eyes reduced,<br />

moderately small, slightly convex. Tempora convex, very<br />

long (about as long as eyes). Mentum: medial tooth acute<br />

apically, slightly shorter than lateral lobes. Ligula narrow<br />

apically. Paraglossae membranous, prominent, much<br />

longer than ligula. Palpi: terminal segment fusiform, not<br />

elliptical, sparsely setulose; penultimate maxillary segment<br />

sparsely setulose. Thorax. Pronotum (Fig. 138) moderately<br />

convex, strongly punctate and wrinkled throughout,<br />

very transverse, moderately cordate; apex straight; anterolateral<br />

angles poorly developed, obtuse; sides strongly<br />

rounded anteriorly, slightly sinuate posteriorly; lateral<br />

beads narrow; lateral depressions absent; two setiferous<br />

punctures on each side; each anterolateral setiferous<br />

puncture situated well in front <strong>of</strong> middle and touching<br />

lateral bead; posterolateral angles obtuse, denticulate;<br />

laterobasal foveae absent; posterior bead interrupted<br />

medially; base emarginate medially, moderately oblique<br />

laterally, slightly narrower than apex, much narrower than<br />

elytral base. Prosternum punctate, wrinkled. Proepisterna<br />

punctate, wrinkled. Metepisterna impunctate. Elytra.<br />

Depressed dorsally, slightly convex laterally, subovate,<br />

widest about middle. Basal margin complete. Shoulders<br />

obtuse, denticulate. Sides moderately rounded. Scutellar<br />

setiferous pore inserted at junction <strong>of</strong> striae 1+2. Scutellar<br />

striole impunctate. Striae complete, moderately deep<br />

(stria 7 shallower basally), finely punctate; stria 3 without<br />

setiferous punctures. Intervals depressed. Sutural apices<br />

angular. Abdomen. Sterna coarsely punctate, wrinkled.<br />

Last visible sternum (sternum VII): male with two apical<br />

ambulatory setae; female with four apical and two medial<br />

ambulatory setae. Aedeagus. Lateral view (Fig. 90):<br />

slightly arcuate; apex rather moderately wide, subtruncate,

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