Fauna of New Zealand 69 - Landcare Research

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Subfamily TRECHINAE
Tribe MORIOMORPHINI
Description (New Zealand). Body length, 3.2–10.3 mm.
Pedunculate or not. Head. Mandibles short to very long,
with setiferous puncture in scrobe. Labrum truncate to
strongly emarginate anteriorly. Clypeus with a setiferous
puncture on each side. Antennae filiform or submoniliform;
segments 1–3 usually glabrous (excluding apical setae);
segment 2 densely pubescent in apical third (Selenochilus,
Trichopsida simplex), with verticillate setae medially (Selenochilus);
segment 3 densely pubescent (in apical third
in Selenochilus; entirely in Trichopsida simplex). Eyes
present; usually two supraorbital setiferous punctures on
inner side, rarely with a single posterior puncture. Tempora
inflated or not. Mentum tooth present medially, entire.
Mentum-submentum suture present. Submentum with four
setae. Ligula apically truncate, arcuate or conical, narrow
or broad, bisetose. Palpi: terminal segment fusiform, rarely
elliptical, glabrous or setulose; penultimate maxillary
segment glabrous or setulose. Thorax. Pronotum usually
with two setiferous punctures on each side (sometimes
with a single anterior puncture, rarely without puncture).
Scutellum entirely or partly visible, elongate or broad,
inserted either partly between and above elytral bases, or
entirely between elytral bases. Legs. Tarsi usually glabrous
dorsally (pubescent in Selenochilus); segments 1–3 of
male protarsi dilated and biseriately pubescent ventrally;
segment 4 of pro- and mesotarsi of both sexes bilobed
(Rossjoycea, Tarastethus) or emarginate; claws entire
ventrally; unguitractor plate invisible between tarsal claws.
Elytra. Usually fused along suture (hindwings vestigial),
rarely free (hindwings fully developed; Mecyclothorax,
in part). Stria 1 not recurrent apically. Discal setiferous
punctures present or absent. Interval 8 carinate or not apically.
Umbilicate series present, separated into two major
groups (usually 7+6 setiferous punctures, sometimes 5+5
punctures, rarely 8+6 punctures). Radial field without
fine dense pubescence. Sutural apices angular to rounded.
Epipleura twisted (with inner fold or plica) near apex. Abdomen.
Apex invisible dorsally. Sterna IV–VI with paired
ambulatory setae only; last visible sternum (sternum VII)
of male usually with two or four apical ambulatory setae,
rarely with six setae; last visible sternum (sternum VII) of
female usually with four apical ambulatory setae (rarely
with six or eight setae) and usually with two medial setae
(rarely with four setae). Aedeagus (Fig. 61–92). Base
closed, bulbose. Parameres. Dissimilar in shape (right
paramere smaller; left paramere elongate or conchiform,
broader basally), glabrous or setulose.
Larochelle & Larivière (2013): Carabidae (Insecta: Coleoptera)
References. Larivière & Larochelle, 2007a: 36–38, (as
Mecyclothoracini, Meonini, Tropopterini; description).
Liebherr, 2011a: 20–21 (description, higher classification),
2011b: 280–300 (higher classification, key to New Zealand
subtribes and genera).
Remarks. Liebherr (2011a) recently synonymised the
tribes Amblytelini, Melisoderini, Tropopterini, Mecyclothoracini,
and Meonini under the tribe Moriomorphini.
The genera Mecyclothorax, Meonochilus (Amblytelina),
and Rossjoycea (Moriomorphina) have been recently
revised (Liebherr & Marris, 2009; Liebherr, 2011b); an
alphabetical checklist of taxa with habitus photos and
species distribution maps is provided here. A taxonomic
revision of the genera Selenochilus (Amblytelina), Molopsida,
Tarastethus (reinstated), and Trichopsida new
genus (Moriomorphina), is presented below.
Key to genera of Moriomorphini
1 Elytra with interval 8 carinate apically (Fig. 19) ....... 2
—Elytra with interval 8 not carinate apically (Fig. 20) 5
2(1) Terminal segment of palpi setulose (Fig. 21). Posterior
bead of pronotum present (complete or interrupted
medially; Fig. 23–24) ....................................
..........................…(p. 62)… Trichopsida new genus
─Terminal segment of palpi glabrous (Fig. 22). Posterior
bead of pronotum absent (Fig. 25) ......................... 3
3(2) Segment 4 of pro- and mesotarsi of both sexes emarginate
apically (cleft for less than half their length;
Fig. 26–27) .......................(p. 48)... Molopsida White
─ Segment 4 of pro- and mesotarsi of both sexes bilobed
apically (cleft for more than half their length; Fig.
28) .......................................................................... 4
4(3) Elytra: stria 3 with three to five setiferous punctures;
stria 5 with one or two setiferous punctures. Segment
5 of metatarsi with four pairs of ventral setae (Fig.
29). [Body length over 9.0 mm.] ..............................
...............................…(p. 55)… Rossjoycea Liebherr
─Elytra: striae 3 and 5 without setiferous punctures. Segment
5 of metatarsi with a single pair of ventral setae
(Fig. 30). [Body length under 7.0 mm.] .....................
.................................. …(p. 56)… Tarastethus Sharp
5(1) Tarsi pubescent dorsally (Fig. 31). Labrum strongly
emarginate (Fig. 33) anteriorly; anterior marginal
setae not equidistant (four setae grouped medially).
Antennae (Fig. 36): segment 2 densely pubescent in
apical third, with verticillate setae medially; segment
3 densely pubescent in apical third ...........................
...........................…(p. 42)… Selenochilus Chaudoir