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Fauna of New Zealand 69 - Landcare Research

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<strong>Fauna</strong> <strong>of</strong> <strong>New</strong> <strong>Zealand</strong> <strong>69</strong> 53<br />

Description. Body: length 5.3–6.0 mm. Head, pronotum,<br />

elytra, and abdomen black; antennae, palpi, and legs<br />

entirely pale red. Microsculpture absent on head and pronotum,<br />

strong and very transverse (with microlines) on<br />

elytra. Iridescence absent on head and pronotum, strong on<br />

elytra (as in M. polita). Very shiny on head and pronotum,<br />

moderately shiny on elytra. Head. Coarsely punctate in<br />

and between frontal furrows, unwrinkled dorsally, much<br />

narrower across eyes than pronotal apex. Mandibles<br />

moderately long. Labrum quadrate, subtruncate anteriorly.<br />

Antennae subfiliform; segment 1 (scape) moderately long,<br />

about 1.5× longer than its maximum width. Frontal furrows<br />

wide, deep. Eyes normally developed, moderately large,<br />

moderately convex; two supraorbital setiferous punctures<br />

on inner side <strong>of</strong> each eye. Interocular fovea present,<br />

deep, rounded. Tempora inflated, oblique, short (about<br />

one-third as long as eyes). Mentum: medial tooth acute<br />

apically, moderately shorter than lateral lobes. Paraglossae<br />

membranous, prominent, much longer than ligula. Palpi:<br />

penultimate maxillary segment glabrous. Thorax. Pronotum<br />

(Fig. 113) strongly convex, coarsely punctate across<br />

base, unwrinkled, moderately transverse, subrectangular;<br />

apex straight; anterior bead narrowly interrupted medially;<br />

anterolateral angles poorly developed, obtusely rounded;<br />

sides moderately rounded anteriorly, very slightly sinuate<br />

posteriorly; lateral beads slightly widening from apex to<br />

base; lateral depressions absent; two setiferous punctures<br />

on each side; each anterolateral setiferous puncture situated<br />

well in front <strong>of</strong> middle and touching lateral bead;<br />

posterolateral angles subrectangular, not denticulate;<br />

laterobasal foveae ill-defined, coarsely punctate, shallow,<br />

narrow, vaguely elongate; base rather straight, much wider<br />

than apex, slightly narrower than elytral base. Prosternum<br />

punctate, unwrinkled. Proepisterna punctate, unwrinkled.<br />

Metepisterna punctate. Legs. Segment 5 <strong>of</strong> metatarsi with a<br />

single pair <strong>of</strong> ventral setae. Elytra. Strongly convex, ovate,<br />

widest in front <strong>of</strong> middle. Shoulders obtuse, denticulate.<br />

Sides moderately rounded. Scutellar striole usually absent,<br />

when present bipunctate. Striae mostly complete (stria 7<br />

obsolete basally), moderately deep, with moderately coarse<br />

punctures. Intervals depressed, becoming slightly convex<br />

apically. Sutural apices angular. Abdomen. Sterna coarsely<br />

punctate, unwrinkled. Last visible sternum (sternum VII):<br />

male with two apical ambulatory setae; female with four<br />

apical and two medial ambulatory setae. Aedeagus. Lateral<br />

view (Fig. 68): moderately arcuate; apex moderately wide,<br />

strongly rounded, moderately projecting ventrally, notched<br />

dorsally. Dorsal view: asymmetrical (ostium deflected to<br />

the right); apex rather straight. Parameres. Setulose (with<br />

a few short setae at apex).<br />

Material examined. 85 specimens (AMNZ, BMNH,<br />

CMNZ, JNNZ, LUNZ, MONZ, NZAC).<br />

Geographic distribution (Map p. 176). North Island: BP,<br />

GB, HB, RI, TK, TO, WA, WI, WN, WO. South Island:<br />

KA, SD<br />

Ecology. Lowland, montane, subalpine. Wet forests<br />

(beech, broadleaf, podocarp) and shrublands. Shaded<br />

ground; wet soil. Nocturnal; hides during the day in and<br />

under fallen rotten branches as well as in and under rotten<br />

logs. The species is xylophilous (associated with wood).<br />

Biology. Seasonality: September to August. Tenerals:<br />

October, December to March. Predacious (based on<br />

mouthpart morphology). Occasionally infested with fungi<br />

(Laboulbeniales).<br />

Dispersal power. Elytra fused basally along suture.<br />

Subapterous. Moderate runner. Vagility limited by flight<br />

incapacity.<br />

Reference. Larochelle & Larivière, 2001: 68–<strong>69</strong> (catalogue;<br />

biology, dispersal power, ecology, geographic<br />

distribution, references).<br />

Remarks. Examination <strong>of</strong> the type <strong>of</strong> Tarastethus dubius<br />

revealed it to be conspecific with Molopsida strenua. As<br />

in the case <strong>of</strong> M. polita, M. strenua is a highly distinctive<br />

species characterised by strongly iridescent elytra.<br />

Molopsida cordipennis (Broun, 1912) E<br />

Figures <strong>69</strong>, 114, 209; Map p. 176<br />

Tarastethus cordipennis Broun, 1912: 388. Holotype: female<br />

(BMNH) labelled “Type (circular red-bordered label; typed)<br />

/ 3173. (hand-written) / <strong>New</strong> Zeal. Broun Coll. Brit. Mus.<br />

1922–482. (white label with red horizontal line; typed) /<br />

Mt. Quoin. Tararua. (hand-written) / Tarastethus cordipennis<br />

(hand-written).” Condition: Excellent.<br />

Molopsida cordipennis: Britton, 1940: 277.<br />

Description. Body: length 5.0–6.0 mm. Head, pronotum,<br />

and elytra testaceous; antennae, palpi, and legs yellowish<br />

red. Microsculpture absent. Iridescence absent. Very shiny.<br />

Head. Coarsely punctate in and between frontal furrows,<br />

unwrinkled dorsally, slightly narrower across eyes than<br />

pronotal apex. Mandibles very long. Labrum slightly<br />

transverse, truncate anteriorly. Antennae submoniliform;<br />

segment 1 (scape) moderately long, about 1.5× longer<br />

than its maximum width. Frontal furrows wide, deep. Eyes<br />

reduced, very small, slightly convex; two supraorbital<br />

setiferous punctures on inner side <strong>of</strong> each eye. Interocular<br />

fovea absent. Tempora inflated, oblique, very long (about<br />

as long as eyes). Mentum: medial tooth acute apically,<br />

slightly shorter than lateral lobes. Paraglossae membranous,<br />

prominent, much longer than ligula. Palpi: penultimate<br />

maxillary segment barely setulose (with a single short<br />

apical seta). Thorax. Pronotum (Fig. 114) strongly convex,<br />

coarsely punctate across base, finely punctate along lateral<br />

beads, wrinkled, elongate, moderately cordate; apex

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