Fauna of New Zealand 69 - Landcare Research

Fauna of New Zealand 69 73
unwrinkled. Last visible sternum (sternum VII): male with
two apical ambulatory setae; female with four apical and
two or four medial ambulatory setae. Aedeagus. Lateral
view (Fig. 86): strongly arcuate; apex narrow, elongate,
rounded at tip, not projecting dorsoventrally. Dorsal view:
asymmetrical (ostium deflected to the right); apex deflected
to the right. Parameres. Glabrous.
Material examined. Four specimens (NZAC).
Geographic distribution (Map p. 182). South Island:
NN─Paturau (Twin Forks Cave area).
Ecology. Lowland. Wet forest (beech) area. Shaded
ground; wet soil. Nocturnal; hides during the day under
logs.
Biology. Seasonality: December, January. Predacious
(based on mouthpart morphology). One specimen infested
by fungi (Laboulbeniales).
Dispersal power. Elytra fused basally along suture.
Subapterous. Moderate runner. Vagility limited by flight
incapacity.
Remarks. This species is named after the type locality,
Paturau (NN), and the Latin suffix -ensis, denoting a place,
locality, or country (Brown, 1985).
Trichopsida paturauensis is morphologically close to
T. pretiosa from which it can be most reliably diagnosed
by the male genitalia, especially the narrow, rounded
apex of the aedeagus. In addition, T. paturauensis has the
following distinguishing features: antennae partially pale
yellowish, with segments 4–6 infuscate; elytra mostly
black, with striae 6–7 obsolete basally.
Trichopsida koyai E new species
Figures 87, 133, 228; Map p. 182
Trichopsida koyai Larochelle & Larivière, new species. Holotype:
male (NZAC) labelled “Mt Stokes Ridge, 2,500’
Marlborough (hand-written) / Attacked by fungoid growth.
(hand-written) / Coll. A.C. O’Connor Oct. 1944 (hand-written)
/ A.E. Brookes Collection (typed) / HOLOTYPE [male
symbol] Trichopsida koyai Larochelle & Larivière, 2013
(red label; typed).” Paratype: one female (NZAC) from the
same locality as the holotype, bearing blue paratype label.
Description. Body: length 4.3–5.4 mm. Head, pronotum,
and elytra mostly blackish brown; abdomen dark brown;
antennae, palpi, and tarsi pale red; femora and tibiae
blackish brown. Microsculpture very transverse (with
microlines), feeble on head and pronotum, strong on
elytra. Iridescence absent. Very shiny. Head. Finely and
sparsely punctate in frontal furrows, unwrinkled dorsally,
slightly narrower across eyes than pronotal apex. Mandibles
moderately long. Labrum moderately transverse
and emarginate anteriorly. Antennae: segment 1 (scape)
moderately long, about 1.5× longer than its maximum
width; segments 1–3 glabrous (excluding apical setae).
Frontal furrows narrow, deep. Eyes reduced, very small,
very slightly convex. Tempora convex, very long (about
as long as eyes). Mentum: medial tooth acute apically,
slightly shorter than lateral lobes. Ligula narrow apically.
Paraglossae membranous, prominent, much longer than
ligula. Palpi: terminal segment fusiform, not elliptical,
sparsely setulose; penultimate maxillary segment sparsely
setulose. Thorax. Pronotum (Fig. 133) moderately convex,
coarsely and sparsely punctate across base, unwrinkled
or sparsely wrinkled along median longitudinal impression,
moderately transverse, subquadrate; apex straight;
anterolateral angles poorly developed, obtusely rounded;
sides moderately rounded anteriorly, not sinuate posteriorly;
lateral beads narrow; lateral depressions absent;
two setiferous punctures on each side; each anterolateral
setiferous puncture situated well in front of middle and
touching lateral bead; posterolateral angles obtuse,
denticulate; laterobasal foveae ill-defined, very shallow,
narrow, round; posterior bead interrupted medially; base
emarginate medially, moderately oblique laterally, slightly
narrower than apex, much narrower than elytral base.
Prosternum punctate, unwrinkled. Proepisterna punctate,
unwrinkled. Metepisterna impunctate. Elytra. Depressed
dorsally, slightly convex laterally, subovate, widest about
middle. Basal margin complete. Shoulders obtuse, not
denticulate. Sides slightly rounded. Scutellar setiferous
pore inserted at base of stria 1 or at junction of striae
1+2. Scutellar striole impunctate. Striae mostly complete
(stria 7 obsolete basally), moderately deep, very finely
punctate (almost impunctate); stria 3 with three setiferous
punctures. Intervals slightly convex, becoming moderately
convex apically. Sutural apices angular. Abdomen. Sterna
impunctate, unwrinkled. Last visible sternum (sternum
VII): male with two apical ambulatory setae; female with
four apical and two medial ambulatory setae. Aedeagus.
Lateral view (Fig. 87): slightly arcuate; apex narrow, subtriangular,
rounded at tip, not projecting dorsoventrally.
Dorsal view: asymmetrical (ostium deflected to the right);
apex deflected to the right. Parameres. Glabrous.
Material examined. Four specimens (NZAC).
Geographic distribution (Map p. 182). South Island:
SD–Mount Robinson. Mount Stokes.
Ecology. Lowland (hill), montane. Forest (beech) area.
Probably shaded ground. Nocturnal; hiding during the
day in leaf litter and under a stone. The body shape
(poorly-developed eyes, inflated tempora, depressed
elytra) suggests that the species is endogean (living in
soil crevices or fissures, in deep humus of leaf litter, under
well-embedded stones).
Biology. Seasonality: October, March. Predacious (based
on mouthpart morphology). Occasionally infested by fungi
(Laboulbeniales).