Fauna of New Zealand 69 - Landcare Research
Fauna of New Zealand 69 - Landcare Research
Fauna of New Zealand 69 - Landcare Research
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<strong>Fauna</strong> <strong>of</strong> <strong>New</strong> <strong>Zealand</strong> <strong>69</strong> 65<br />
(scape) moderately long, about 1.5× longer than its maximum<br />
width, glabrous (excluding apical setae); segment<br />
2 densely pubescent in apical third (glabrous, excluding<br />
apical setae, in other species); segment 3 densely pubescent<br />
(glabrous, excluding apical setae, in other species).<br />
Frontal furrows wide, shallow. Eyes normally developed,<br />
moderately large, strongly convex. Tempora oblique, short<br />
(about one-third as long as eyes). Mentum: medial tooth<br />
acute apically, slightly shorter than lateral lobes. Ligula<br />
narrow apically. Paraglossae membranous, prominent,<br />
much longer than ligula. Palpi: terminal segment fusiform,<br />
not elliptical, sparsely setulose; penultimate maxillary<br />
segment sparsely setulose. Thorax. Pronotum (Fig. 123)<br />
moderately convex, impunctate, feebly wrinkled basally<br />
as well as along median longitudinal impression and apex,<br />
strongly transverse, trapezoid; apex strongly arcuate; anterolateral<br />
angles strongly developed, obtusely rounded;<br />
sides strongly rounded anteriorly, not sinuate posteriorly;<br />
lateral beads narrow; lateral depressions absent; two setiferous<br />
punctures on each side; each anterolateral setiferous<br />
puncture situated well in front <strong>of</strong> middle and touching<br />
lateral bead; posterolateral angles acute and projecting<br />
laterally, denticulate; laterobasal foveae absent; posterior<br />
bead interrupted medially; base emarginate medially,<br />
slightly oblique laterally, slightly narrower than apex,<br />
much wider than elytral base. Prosternum impunctate,<br />
unwrinkled. Proepisterna impunctate, unwrinkled. Metepisterna<br />
impunctate. Elytra. Moderately convex, ovate,<br />
widest about middle. Basal margin incomplete, only<br />
reaching level <strong>of</strong> stria 3 (complete, reaching level <strong>of</strong> stria<br />
1 in other species). Shoulders rounded, denticulate. Sides<br />
strongly rounded. Scutellar setiferous pore absent. Scutellar<br />
striole bipunctate. Striae complete, shallow (striae<br />
5–7 shallower), virtually impunctate; stria 3 with a single<br />
setiferous puncture (behind middle). Intervals depressed,<br />
becoming slightly convex apically. Sutural apices angular.<br />
Abdomen. Sterna impunctate, unwrinkled. Last visible<br />
sternum (sternum VII): male with six apical ambulatory<br />
setae (with two or four setae in other species); female with<br />
six apical and two medial ambulatory setae. Aedeagus.<br />
Lateral view (Fig. 78): strongly arcuate; apex narrow,<br />
subtriangular-rounded, strongly projecting ventrally<br />
(more so than in other species). Dorsal view: asymmetrical<br />
(ostium deflected to the right); apex deflected to the<br />
right. Parameres. Left paramere glabrous; right paramere<br />
setulose (with numerous short ventral setae in apical half).<br />
Material examined. 31 specimens (BMNH, CMNZ,<br />
JNNZ, NZAC).<br />
Geographic distribution (Map p. 183). South Island:<br />
DN–Allison Scenic Reserve (near Akatore). Measly Beach<br />
(near Waikaro). Port Chalmers. Taieri Mouth (Picnic<br />
Gully). Waipori Falls. Waipori Gorge. Waipori Valley<br />
(Government Track). SL–Tapanui (Whisky Gully).<br />
Ecology. Lowland. Wet forests (broadleaved, podocarp,<br />
beech). Shaded ground; wet soil. Nocturnal; hides during<br />
the day mostly in the soil and under stones, sometimes<br />
in leaf litter. The species is truly endogean (living in soil<br />
crevices or fissures, in deep humus <strong>of</strong> leaf litter, under<br />
well-embedded stones). It is best collected by soil-washing<br />
techniques.<br />
Biology. Seasonality: September, December to February,<br />
April to June. Tenerals: May. Predacious (based on<br />
mouthpart morphology).<br />
Dispersal power. Elytra fused basally along suture.<br />
Subapterous. Moderate runner. Vagility limited by flight<br />
incapacity.<br />
Reference. Larochelle & Larivière, 2001: 68 (biology,<br />
dispersal power, ecology, geographic distribution).<br />
Remarks. Trichopsida simplex differs from all other<br />
Trichopsida species by the following characters: antennae<br />
with segment 2 densely pubescent in apical third (segment<br />
2 glabrous, excluding apical setae in other species), segment<br />
3 densely pubescent (segment 3 glabrous excluding<br />
apical setae in other species); elytra with basal margin<br />
incomplete, only reaching level <strong>of</strong> stria 3 (complete,<br />
reaching level <strong>of</strong> stria 1 in other species); last visible<br />
sternum (sternum VII) <strong>of</strong> male abdomen with six apical<br />
ambulatory setae (with two or four setae in other species);<br />
apex <strong>of</strong> aedeagus strongly projecting ventrally (more so<br />
than in other species). The above-mentioned characters<br />
are generally indicative <strong>of</strong> species-level variation among<br />
Moriomorphina and probably do not warrant a different<br />
subgeneric or generic status for this species. Trichopsida<br />
simplex simply appears to be an odd-looking species <strong>of</strong><br />
Trichopsida.<br />
Trichopsida nitida E new species<br />
Figures 124, 219; Map p. 182<br />
Trichopsida nitida Larochelle & Larivière, new species. Holotype:<br />
female (NZAC) labelled “NEW ZEALAND BR<br />
Tiropahi River 140m 11 Feb 1986 J. Nunn & J.I. Townsend<br />
(typed) / NZMS 260: K30: 810144 (typed) / HOLOTYPE<br />
[female symbol] Trichopsida nitida Larochelle & Larivière,<br />
2013 (red label; typed).”<br />
Description. Body: length 7.6 mm. Head, pronotum, and<br />
elytra black; lateral margins and apex <strong>of</strong> elytra yellowish<br />
red; abdomen black; antennae, palpi, and legs entirely<br />
pale red. Microsculpture absent. Iridescence absent. Very<br />
shiny. Head. Coarsely punctate in frontal furrows, wrinkled<br />
on frons, slightly narrower across eyes than pronotal<br />
apex. Mandibles short. Labrum moderately transverse,<br />
asymmetrical, slightly emarginate anteriorly. Antennae:<br />
segment 1 (scape) moderately long, about 2× longer than<br />
its maximum width; segments 1–3 glabrous (excluding<br />
apical setae). Frontal furrows wide, shallow. Eyes normally