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Fauna of New Zealand 69 - Landcare Research

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<strong>Fauna</strong> <strong>of</strong> <strong>New</strong> <strong>Zealand</strong> <strong>69</strong> 65<br />

(scape) moderately long, about 1.5× longer than its maximum<br />

width, glabrous (excluding apical setae); segment<br />

2 densely pubescent in apical third (glabrous, excluding<br />

apical setae, in other species); segment 3 densely pubescent<br />

(glabrous, excluding apical setae, in other species).<br />

Frontal furrows wide, shallow. Eyes normally developed,<br />

moderately large, strongly convex. Tempora oblique, short<br />

(about one-third as long as eyes). Mentum: medial tooth<br />

acute apically, slightly shorter than lateral lobes. Ligula<br />

narrow apically. Paraglossae membranous, prominent,<br />

much longer than ligula. Palpi: terminal segment fusiform,<br />

not elliptical, sparsely setulose; penultimate maxillary<br />

segment sparsely setulose. Thorax. Pronotum (Fig. 123)<br />

moderately convex, impunctate, feebly wrinkled basally<br />

as well as along median longitudinal impression and apex,<br />

strongly transverse, trapezoid; apex strongly arcuate; anterolateral<br />

angles strongly developed, obtusely rounded;<br />

sides strongly rounded anteriorly, not sinuate posteriorly;<br />

lateral beads narrow; lateral depressions absent; two setiferous<br />

punctures on each side; each anterolateral setiferous<br />

puncture situated well in front <strong>of</strong> middle and touching<br />

lateral bead; posterolateral angles acute and projecting<br />

laterally, denticulate; laterobasal foveae absent; posterior<br />

bead interrupted medially; base emarginate medially,<br />

slightly oblique laterally, slightly narrower than apex,<br />

much wider than elytral base. Prosternum impunctate,<br />

unwrinkled. Proepisterna impunctate, unwrinkled. Metepisterna<br />

impunctate. Elytra. Moderately convex, ovate,<br />

widest about middle. Basal margin incomplete, only<br />

reaching level <strong>of</strong> stria 3 (complete, reaching level <strong>of</strong> stria<br />

1 in other species). Shoulders rounded, denticulate. Sides<br />

strongly rounded. Scutellar setiferous pore absent. Scutellar<br />

striole bipunctate. Striae complete, shallow (striae<br />

5–7 shallower), virtually impunctate; stria 3 with a single<br />

setiferous puncture (behind middle). Intervals depressed,<br />

becoming slightly convex apically. Sutural apices angular.<br />

Abdomen. Sterna impunctate, unwrinkled. Last visible<br />

sternum (sternum VII): male with six apical ambulatory<br />

setae (with two or four setae in other species); female with<br />

six apical and two medial ambulatory setae. Aedeagus.<br />

Lateral view (Fig. 78): strongly arcuate; apex narrow,<br />

subtriangular-rounded, strongly projecting ventrally<br />

(more so than in other species). Dorsal view: asymmetrical<br />

(ostium deflected to the right); apex deflected to the<br />

right. Parameres. Left paramere glabrous; right paramere<br />

setulose (with numerous short ventral setae in apical half).<br />

Material examined. 31 specimens (BMNH, CMNZ,<br />

JNNZ, NZAC).<br />

Geographic distribution (Map p. 183). South Island:<br />

DN–Allison Scenic Reserve (near Akatore). Measly Beach<br />

(near Waikaro). Port Chalmers. Taieri Mouth (Picnic<br />

Gully). Waipori Falls. Waipori Gorge. Waipori Valley<br />

(Government Track). SL–Tapanui (Whisky Gully).<br />

Ecology. Lowland. Wet forests (broadleaved, podocarp,<br />

beech). Shaded ground; wet soil. Nocturnal; hides during<br />

the day mostly in the soil and under stones, sometimes<br />

in leaf litter. The species is truly endogean (living in soil<br />

crevices or fissures, in deep humus <strong>of</strong> leaf litter, under<br />

well-embedded stones). It is best collected by soil-washing<br />

techniques.<br />

Biology. Seasonality: September, December to February,<br />

April to June. Tenerals: May. Predacious (based on<br />

mouthpart morphology).<br />

Dispersal power. Elytra fused basally along suture.<br />

Subapterous. Moderate runner. Vagility limited by flight<br />

incapacity.<br />

Reference. Larochelle & Larivière, 2001: 68 (biology,<br />

dispersal power, ecology, geographic distribution).<br />

Remarks. Trichopsida simplex differs from all other<br />

Trichopsida species by the following characters: antennae<br />

with segment 2 densely pubescent in apical third (segment<br />

2 glabrous, excluding apical setae in other species), segment<br />

3 densely pubescent (segment 3 glabrous excluding<br />

apical setae in other species); elytra with basal margin<br />

incomplete, only reaching level <strong>of</strong> stria 3 (complete,<br />

reaching level <strong>of</strong> stria 1 in other species); last visible<br />

sternum (sternum VII) <strong>of</strong> male abdomen with six apical<br />

ambulatory setae (with two or four setae in other species);<br />

apex <strong>of</strong> aedeagus strongly projecting ventrally (more so<br />

than in other species). The above-mentioned characters<br />

are generally indicative <strong>of</strong> species-level variation among<br />

Moriomorphina and probably do not warrant a different<br />

subgeneric or generic status for this species. Trichopsida<br />

simplex simply appears to be an odd-looking species <strong>of</strong><br />

Trichopsida.<br />

Trichopsida nitida E new species<br />

Figures 124, 219; Map p. 182<br />

Trichopsida nitida Larochelle & Larivière, new species. Holotype:<br />

female (NZAC) labelled “NEW ZEALAND BR<br />

Tiropahi River 140m 11 Feb 1986 J. Nunn & J.I. Townsend<br />

(typed) / NZMS 260: K30: 810144 (typed) / HOLOTYPE<br />

[female symbol] Trichopsida nitida Larochelle & Larivière,<br />

2013 (red label; typed).”<br />

Description. Body: length 7.6 mm. Head, pronotum, and<br />

elytra black; lateral margins and apex <strong>of</strong> elytra yellowish<br />

red; abdomen black; antennae, palpi, and legs entirely<br />

pale red. Microsculpture absent. Iridescence absent. Very<br />

shiny. Head. Coarsely punctate in frontal furrows, wrinkled<br />

on frons, slightly narrower across eyes than pronotal<br />

apex. Mandibles short. Labrum moderately transverse,<br />

asymmetrical, slightly emarginate anteriorly. Antennae:<br />

segment 1 (scape) moderately long, about 2× longer than<br />

its maximum width; segments 1–3 glabrous (excluding<br />

apical setae). Frontal furrows wide, shallow. Eyes normally

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