GENE TIC REPROD UCTION. 9 1and the quiescent sporocyst states, the first yieldingthe essential andspecific form, are now found to obtain throughout the majorityof theFlagellata. Simultaneously with, and in many instances anterior to, thepublication of Messrs. Dallinger and Drysdale's researches, correspondingphenomena had been observed and duly recorded by the present author inrelation with the representatives of the Choano-Flagellata or collarbearingsection, and other more simply organized Flagellata. Like resultshave also accompanied his more recent investigations, while confirmatoryevidence in the same direction isabundantly afforded in the magnificentvolume devoted to the illustration of the Flagellata generally, lately publishedby Professor Stein. As examples of the persistence of the threeabove-named characteristic phases in the group of the Choano-Flagellata,referencemay be more especially made to the systematic descriptions andaccompanying figures of such species as Salpingceca amphoridium, S. fusiformis,and Codosiga botrytis, in all of which the transitory amoeboid phaseis particularly remarkable. In all those Eustomatous Flagellata which,like Phacus and Anisonema, possess an indurated cuticle, the assumptionof an amoeboid condition would not be possible, and the animalcule previousto encystment and spore development merely loses its flagella, andassumes a quiescent state. In Euglena and its allies, including Eutreptia,the animalcules exhibit a transitory amcebiform, or, more correctly, agregarine-like repent state immediately preceding the process of encystment.With the representatives of the two last-named generait is furtherworthy of note that the initial condition of existence on emerging fromthe sporeis likewise amcebiform and non-flagelliferous. Among the Choano-Flagellata this earliest stage is simply flagellate or monadiform, therebeing no trace of the characteristic collar, while in a larger series, includingthe majority of the Pantostomata, such initial phase, in all but size,corresponds essentially with the parent zooid.Sexual or Genetic Reproduction.Sexual reproduction in the broad acceptation of the term as belongingto the propagative phenomena of the higher animals or Metazoa, or, asup to within a comparatively recent date maintained by Balbiani andothers to obtain among the members of the higher Ciliate section of Infusoria,and in either case involving the concourse of true and independentlydeveloped sexual cellular elements ova and spermatozoa remains, sofar as it concerns all or any members of the class or classes now underdiscussion, entirely undemonstrable. Nor, the essentially unicellular natureor value of the infusorial organism, as here advocated, being once firmlyand incontestably established, can a contrary verdict be anticipated!Notwithstanding, however, the apparently uncompromising verdict pronouncedin the foregoing sentence, it will presently be shown that theseProtozoic organisms, in essence if not in fact, fulfil a r61e of reproduction
92 ORGANIZATION OF THE INFUSORIA.that corresponds most closely with this importantfunction as it is metwith in all Metazoic or multicellular structures. As already maintainedin the description of that more common phase of multiplication whichtakes the form of constantly repeated binary division, the Infusoriathrough such mode of increase merely repeat in a separate and inde-which characterises thependent manner that process of cell-multiplicationnormal growths of all tissue structures. By-and-by, however, in the case ofthe Metazoon or tissue organism, an epochis arrived at when the componentcells cease to exhibit their previous duplicative energy, the consequencebeing the gradual decay and ultimate dissolution or death of theentire organism. But for the interposition of a special and more or lessperiodical regenerative act, this termination of the life of the individualwould also, sooner or later, involve the extinction of the race or species.Such a regenerative act, and the further survival of the race, is, however,here accomplished through the fusion or union of one of that congeries of cellsout of which the compound organism is composed with one other exteriorlyderived cell liberated from another organism or congeries of cells pertainingto the same specific type. The result of such fusion between these two,denominated respectively the germ-cell and sperm-cell, or, in other words,the ovum and spermatic cell or element, is the capacity conferred upon theformer of once more proceeding with the duplicative process, and repeatingthat cycle of cell-aggregation or tissue-construction followed by the parentorganism. In a precisely similar manner, the infusorial body, after repeatedduplicative multiplication, arrives at a condition in which the strain orrace is too exhausted for the further maintenance of this process, andwithout the intervention of some supplementary regenerative operationwould become extinct. By, however, the coalescence or fusion of one ofthese individual cellsor animalcules with an elemental cell or animalculederived from a neighbouring race or strain, the capacity to continue theduplicative process is revived, and the further duration of the race secured.The two animalcules thus uniting with one another correspond to all intentsand purposes with the coalescing germ-cell and sperm-cell, or ovum andspermatozoon of the higher tissue organisms, the only essential point ofdivergence being in the subsequent changes manifested ; the organism inthe latter instance exhibits a tendency to build out of the cells amassedby the duplicative process, more or less complex coherent tissues, while inthe former each cell so produced maintains a separate and independentexistence.Among the extensive series of types included in the infusorial classes, thephenomenon of " conjugation," " fusion," or " zygosis," as it is variously called,and which undoubtedly represents the sexual or genetic reproductive processof the higher animals in its most simplified or elementary form of expression,exhibits certain well-defined modifications. With the majority of the morehighly organized Ciliata, such conjugation has to be denominated, as comparedwith the form next described, transient and incomplete, being, as
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aoamoa
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"Our little systems have their day,
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TOTHOMAS HENRY HUXLEY, LL.D.,F.R.S.
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viiiPREFACE.experience some disappo
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XPREFACE.ready and valuable assista
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LEEUWENHOEtfS OBSERVATIONS. 3relate
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LEEUWENHOEICS OBSERVATIONS.5spatter
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LEEUWENHOEK'S OBSERVATIONS.Jstopped
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SIfi E. KING, 1693. JOHN HARRIS, 16
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STEPHEN GRA Y, 1696. LEEUWENHOEK, 1
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HENRY BAKER, 1742, 1753.13"Oct. 6th
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O. F. MULLER, 1773-1786. 15ledge of
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EHRENBERG, 1836. 17Notwithstanding
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F. DUJARDIN, 1841. T. VON SIEBOLD,
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FRIEDRICH STEIN, 1849-1854. 21cules
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CLAPAREDE AND LACHMANN, 1858-1860.
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F. STEIN, 1859. R. M. DIES ING, 184
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ANDREW PRITCHARD, 1861. H. JAMES-CL
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DALLINGER AND DRYSDALE, 1873-1875.
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CHAPTER II.THE SUB-KINGDOM PROTOZOA
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AFFINITIES OF THE SPONGIDA. 33ordin
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PRIMARY SUBDIVISIONS A UTHOKS S YST
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AUTHORS PHYLOGENETIC SCHEME. 37DIAG
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FLA CELLA TA -PANTOS TOMA TA ; FLA
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A UTHORS INVESTIGA TIONS. 1 4 1clos
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( 143 )CHAPTER V.NATURE AND AFFINIT
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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( 195 )CHAPTER VI.SYSTEMS OF CLASSI
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CLASSIFICATION OF THE INFUSORIA.197
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MULLERS CLA SSIPICA TOR Y S YSTEM.
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EHRENBERG'S CLASSIFICATORY SYSTEM.2
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CLASSIFICATORY SYSTEMS OF SIEBOLD A
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CLAPAREDE AND LACHMANWS CLASSIFICAT
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DIESINGS CLASSIFICATORY SYSTEM. 207
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S TEIN'S CLA SSIFICA TOR Y S Ki TEM
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A UTHOKS CLASSIFICA TOR Y S YSTEM.
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A UTHOR'S CLA SSIPICA TOR Y S YSTEM
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A UTHOKS CLASSIPICA TOR Y S YSTEM.
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CLASS FLAGELLA TA. 2 I7more extensi
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GENUS TRYPANOSOMA. 219Trypanosoma s
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GENUS MASTIGAMCEBA . 221The some ha
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;,HAB.GENUS REPTOMONAS. 22$immediat
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ORDER RA DIO-FLA CELLA TA.225Podost
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Body subspherical orGENUS ACTINOMON
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GENUS SPONGASTERISCUS. 229Spongocyc
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Cladomonas.ipidodendrtSpongomonas.D