UNICELL ULAR NA TURE. 5 3dissolution, is there any appearance of a multicellular constitution, or in theformer instance, more especially, the formation of a distinct germinal layeror blastoderm, the one essential feature and index of all the multicellularanimals or Metazoa. It is true nevertheless, although the circumstancehas apparently as yet attracted but little notice, that many infusorial formsbelonging both to the Ciliate and Flagellate sections of the group, exhibitin connection with that mode of reproduction characterized by the resolutionof the primary cell or zooid into a number of sporular bodies, an aspect andplan of organization in no ways to be distinguished from the moruloid orprimary segmental condition of the ovum of the Metazoa. Beyond thisstage, however, the analogy, or, if it exists, the homology, entirely ceases ;for whereas, in the Infusorium or Protozoon, each segment or unicellularcomponent of the pseudo-morula becomes metamorphosed into a distinctand independent being, in the Metazoon, this primary independence isalmost immediately obliterated through the recasting or reconstruction outof the primary segmental elements, or morula, of the true multicellularembryo with its characteristic inner and outer germinal layers or primordialtissues. In the case of the sponges, as shown later on, and notwithstandingthe deceptiveness of external appearances, the production of similar pseudomorulaeare associated both with the growth of the free-swimming swarmgemmulesor so-called ciliated larvae, and also with the development incertain types of the characteristic ciliated chambers, or, as they are moreusually designated, the ampullaceous sacs of the adult sponge.Taking for granted that all infusorial structures possess a unicellularmorphologic value only, the very extensive range of complexity compatiblewith such simple organization, as exhibited by the representatives ofthe Infusoria, has now to be considered. Among the majorityof theolder biologists, and with many even at the present day, the conceptionof a single histologic cell, or of an independent unicellular organism, differswidely from the one that is here advocated. With the former it was, and is,held, firstly, that such a simple cell or unicellular organism must have adifferentiated bounding membrane, the cell-wall or primordial utricle and;secondly, that the same must contain a central denser and more highlyrefractive mass, or nucleus, which may or may not be associated with a stillmore minute segregated mass, the nucleolus. As demonstrated, however,by the later school of biologists, and among whom Professor Haeckel'sname occupies a pre-eminent position, neither a distinct cell-wall nora differentiated central nucleus forms an essential or invariable elementof cell-organization, be such cell either an independent being, or unit,tissue. In accordanceor an integral constituent only of a compoundwith the results of more modern investigation, the intrinsic value orpotentialityof such a cell-structure resides neither in the cell-wall norin the nucleus, but in the simple protein matter indifferently denominatedsarcode or protoplasm, of which the cell-bodyis built up. Withreference to the more or less highly differentiated organization of cell-
54 ORGANIZATION OF THE INFUSORIA.structures and single-celled organisms, Professor Haeckel has recently proposedto introduce a special code of terminology. In connection with this heconfers upon all those cells, or so-called plastids, in which no nucleus ornucleus-like structure is present, the title of simple cytodes, reserving that oftrue cells for those alone in which such a structure is distinctly represented.Both of these are again recognized by this authority as including two minorgroups of equal value, distinguished by the presence or absence of abounding membrane or cell-wall ;the naked and membrane-boundedcytodes he has denominated respectively gymnocytodes and hullcytodes,and the nucleated cells in a similar manner, Urzellen or gymnocyta, andHullzellen or lepocyta. This separation of the nucleated and non-nucleatedunicellular structures generally, as applied to independent unicellular orProtozoic organisms in particular, forms the basis upon which ProfessorHaeckel has, as previously stated, proposed to establish his non-nucleatedclass-group of the Monera. In recognition of this same distinction, ProfessorHuxley, in his 'Anatomy of the Invertebrata,' has subdivided the Protozoainto the two groups of the Monera and Endoplastica ;the former correspondingwith the group of the same name as established by Haeckel,and the latter including that remaining great majority of the Protozoa inwhich an endoplastic or nucleus-like structure is distinctly visible. Sucha distinction is, nevertheless, adopted by this author as a matter only oftemporary convenience, he freely expressing his doubts as to whether itwill stand the test of extended investigation. The outcome of such researchsince the publicationof Professor Huxley's volume, has indeed fullyjustified the characteristic caution displayed by this eminent biologist ;several of the more important groups of the so-called Monera, includingmore especially the Foraminifera, being now found to consist of nucleatedstructures conforming in all essential details with that larger section of theProtozoa from which it has been proposed to separate them. In accordancewith the opinion maintained by the author of this volume, and as alreadyintimated in the preceding chapter, the Monera, as a distinct class, has nosubstantial claim for retention, all the representatives of the Protozoa beingheld to possess a nucleus, or its equivalent, in their fully matured condition.In their earliest and immature state this important structure, the nucleus,is undoubtedly, however, often absent, the Protozoon, under such conditionsonly, conforming in structure with Professor Haeckel's diagnosisof a simple cytode or Moneron. That a unicellular animal may, on theother hand, be entirely destitute of a differentiated bounding membrane, orcell-wall, is abundantly evident. All such peripheral differentiation isclearly conspicuous for its absence in the whole of that section of theProtozoa here distinguished by the title of the Pantostomata, and in whichfood-substances are incepted indifferently at any point of the periphery.As already indicated in the preceding chapter, this simplest and homogeneoustype of protoplasmic structure, the inseparable corollary of thePantostomatous organism,is found associated with by far the larger portion
- Page 3:
aoamoa
- Page 6 and 7:
"Our little systems have their day,
- Page 9:
TOTHOMAS HENRY HUXLEY, LL.D.,F.R.S.
- Page 12 and 13:
viiiPREFACE.experience some disappo
- Page 14 and 15:
XPREFACE.ready and valuable assista
- Page 16 and 17: LEEUWENHOEtfS OBSERVATIONS. 3relate
- Page 18 and 19: LEEUWENHOEICS OBSERVATIONS.5spatter
- Page 20 and 21: LEEUWENHOEK'S OBSERVATIONS.Jstopped
- Page 22 and 23: SIfi E. KING, 1693. JOHN HARRIS, 16
- Page 24 and 25: STEPHEN GRA Y, 1696. LEEUWENHOEK, 1
- Page 26 and 27: HENRY BAKER, 1742, 1753.13"Oct. 6th
- Page 28 and 29: O. F. MULLER, 1773-1786. 15ledge of
- Page 30 and 31: EHRENBERG, 1836. 17Notwithstanding
- Page 32 and 33: F. DUJARDIN, 1841. T. VON SIEBOLD,
- Page 34 and 35: FRIEDRICH STEIN, 1849-1854. 21cules
- Page 36 and 37: CLAPAREDE AND LACHMANN, 1858-1860.
- Page 38 and 39: F. STEIN, 1859. R. M. DIES ING, 184
- Page 40 and 41: ANDREW PRITCHARD, 1861. H. JAMES-CL
- Page 42 and 43: DALLINGER AND DRYSDALE, 1873-1875.
- Page 44 and 45: CHAPTER II.THE SUB-KINGDOM PROTOZOA
- Page 46 and 47: AFFINITIES OF THE SPONGIDA. 33ordin
- Page 48 and 49: PRIMARY SUBDIVISIONS A UTHOKS S YST
- Page 50 and 51: AUTHORS PHYLOGENETIC SCHEME. 37DIAG
- Page 52 and 53: FLA CELLA TA -PANTOS TOMA TA ; FLA
- Page 54 and 55: CHOANO-FLAGELLATA; MYCETOZOA. 41acc
- Page 56 and 57: MYCETOZOAj LABYRINTHULIDA. 43From t
- Page 58 and 59: GROUPS PROTISTA AND MONERA. 45of th
- Page 60 and 61: DISTINCTION BETWEEN PROTOZOA AND PR
- Page 62 and 63: ( 49 )CHAPTER III.NATURE AND ORGANI
- Page 64 and 65: AUTHORS CLASSIFICATORY TABLE.TABULA
- Page 68 and 69: UN1CELL ULAR NA TURE. 5 5of the ent
- Page 70 and 71: CUTICULAR ELEMENTS. 57substance the
- Page 72 and 73: EXCRETED ELEMENTS. 59by the interca
- Page 74 and 75: EXCRETED ELEMENTS. 6 1transparent,
- Page 76 and 77: ENCYSTMENT. 63corresponding type of
- Page 78 and 79: LOCOMOTIVE AND PREHENSILE APPENDAGE
- Page 80 and 81: ORAL APERTURE. 67Oral Aperture or C
- Page 82 and 83: CONTRACTILE VESICLES. 69shadowed. A
- Page 84 and 85: CONTRACTILE VESICLES.71in the major
- Page 86 and 87: NUCLEUS OR ENDOPLAST. 73to indicate
- Page 88 and 89: NUCLEUS OR ENDOPLAST. 75Spirostomit
- Page 90 and 91: NUCLEOLUS OR ENDOPLASTULE. 77with t
- Page 92 and 93: COLOURING SUBSTANCES. 79held to ind
- Page 94 and 95: TRICHOCYSTS. 8 1crowded together an
- Page 96 and 97: TRICHOCYSTS. 83follow it,and being
- Page 98 and 99: BINARY DIVISION. 85that remain, red
- Page 100 and 101: EXTERNAL GEMMATION. 87entire oblite
- Page 102 and 103: SPORULAR MULTIPLICATION. 89four, ei
- Page 104 and 105: GENE TIC REPROD UCTION. 9 1and the
- Page 106 and 107: GENETIC REPRODUCTION. 93illustrated
- Page 108 and 109: GENETIC REPRODUCTION. 95capsules we
- Page 110 and 111: GENETIC REPRODUCTION. 97be essentia
- Page 112 and 113: ZOOLOGICAL AFFINITIES.90as known, i
- Page 114 and 115: ZOOLOGICAL AFFINITIES.IOIunicellula
- Page 116 and 117:
ZOOLOGICAL AFFINITIES. 103position.
- Page 118 and 119:
ZOOLOGICAL AFFINITIES. 105forming t
- Page 120 and 121:
DIS TRIE UTION. \ O 7plete covering
- Page 122 and 123:
DISTRIBUTION.IOQIn an enumeration o
- Page 124 and 125:
DISTRIBUTION. I Ihunting grounds. A
- Page 126 and 127:
PRESER VA TION. I 13Preservation of
- Page 128 and 129:
ME THODS OF INVES TIGA TION. I I5fo
- Page 130 and 131:
METHODS OF INVESTIGA TION.1 1^only
- Page 132 and 133:
RED I; TUBERVILLE NEEDHAM. 119simil
- Page 134 and 135:
JOHN HARRIS ; SPALLANZANI.l 2 rfor
- Page 136 and 137:
LORENZ OKEN; EHRENBERG. 123moment w
- Page 138 and 139:
POUCHET ; PASTEUR. 125Director of t
- Page 140 and 141:
PROFESSOR TYNDALL. 127now to be sup
- Page 142 and 143:
PROFESSOR TYNDALL. I29needle-dip fr
- Page 144 and 145:
PROFESSOR TYNDALL. 131tubes, as giv
- Page 146 and 147:
DALLINGER AND DRYSDALE. 133four day
- Page 148 and 149:
A UTHOKS INVESTIGA TIONS. 13 5and D
- Page 150 and 151:
A UTHORS INVESTIGA TIONS. 13 7or le
- Page 152 and 153:
AUTHOR'S INVESTIGATIONS. 139and lai
- Page 154 and 155:
A UTHORS INVESTIGA TIONS. 1 4 1clos
- Page 156 and 157:
( 143 )CHAPTER V.NATURE AND AFFINIT
- Page 158 and 159:
NATURE AND AFFINITIES OF THE SPONGE
- Page 160 and 161:
NATURE AND AFFINITIES OF THE SPONGE
- Page 162 and 163:
NATURE AND AFFINITIES OF THE SPONGE
- Page 164 and 165:
NATURE AND AFFINITIES OF THE SPONGE
- Page 166 and 167:
NATURE AND AFFINITIES OF THE SPONGE
- Page 168 and 169:
NATURE AND AFFINITIES OF THE SPONGE
- Page 170 and 171:
NATURE AND AFFINITIES OF THE SPONGE
- Page 172 and 173:
NATURE AND AFFINITIES OF THE SPONGE
- Page 174 and 175:
NATURE AND AFFINITIES OF THE SPONGE
- Page 176 and 177:
NATURE AND AFFINITIES OF THE SPONGE
- Page 178 and 179:
NATURE AND AFFINITIES OF THE SPONGE
- Page 180 and 181:
NATURE AND AFFINITIES OF THE SPONGE
- Page 182 and 183:
NATURE AND AFFINITIES OF THE SPONGE
- Page 184 and 185:
NATURE AND AFFINITIES OF THE SPONGE
- Page 186 and 187:
NATURE AND AFFINITIES OF THE SPONGE
- Page 188 and 189:
NATURE AND AFFINITIES OF THE SPONGE
- Page 190 and 191:
NATURE AND AFFINITIES OF THE SPONGE
- Page 192 and 193:
NATURE AND AFFINITIES OF THE SPONGE
- Page 194 and 195:
NATURE AND AFFINITIES OF THE SPONGE
- Page 196 and 197:
NATURE AND AFFINITIES OF THE SPONGE
- Page 198 and 199:
NATURE AND AFFINITIES OF THE SPONGE
- Page 200 and 201:
NATURE AND AFFINITIES OF THE SPONGE
- Page 202 and 203:
NATURE AND AFFINITIES OF THE SPONGE
- Page 204 and 205:
NATURE AND AFFINITIES OF THE SPONGE
- Page 206 and 207:
NATURE AND AFFINITIES OF THE SPONGE
- Page 208 and 209:
( 195 )CHAPTER VI.SYSTEMS OF CLASSI
- Page 210 and 211:
CLASSIFICATION OF THE INFUSORIA.197
- Page 212 and 213:
MULLERS CLA SSIPICA TOR Y S YSTEM.
- Page 214 and 215:
EHRENBERG'S CLASSIFICATORY SYSTEM.2
- Page 216 and 217:
CLASSIFICATORY SYSTEMS OF SIEBOLD A
- Page 218 and 219:
CLAPAREDE AND LACHMANWS CLASSIFICAT
- Page 220 and 221:
DIESINGS CLASSIFICATORY SYSTEM. 207
- Page 222 and 223:
S TEIN'S CLA SSIFICA TOR Y S Ki TEM
- Page 224 and 225:
A UTHOKS CLASSIFICA TOR Y S YSTEM.
- Page 226 and 227:
A UTHOR'S CLA SSIPICA TOR Y S YSTEM
- Page 228 and 229:
A UTHOKS CLASSIPICA TOR Y S YSTEM.
- Page 230 and 231:
CLASS FLAGELLA TA. 2 I7more extensi
- Page 232 and 233:
GENUS TRYPANOSOMA. 219Trypanosoma s
- Page 234 and 235:
GENUS MASTIGAMCEBA . 221The some ha
- Page 236 and 237:
;,HAB.GENUS REPTOMONAS. 22$immediat
- Page 238 and 239:
ORDER RA DIO-FLA CELLA TA.225Podost
- Page 240 and 241:
Body subspherical orGENUS ACTINOMON
- Page 242 and 243:
GENUS SPONGASTERISCUS. 229Spongocyc
- Page 244 and 245:
Cladomonas.ipidodendrtSpongomonas.D