A UTHOKS INVESTIGA TIONS. 13 5and Drysdale may also be cavilled at as deficient of the necessary proofs,inasmuch as the various species with which their researches are connectedare peculiar to certain putrefying animal macerations only, and cannottherefore be said to fairly represent those types common to organic infusiongenerally, with which the phenomena of de novo generation have been heldmore especially to obtain.Now, among these, infusion of hayhas from the earliest date of thediscovery of Infusoria, and the promulgation of the doctrine of spontaneousgeneration, been recognized as the most productive material for the artificialdevelopment of these minute beings, and as the one in which such presumedde novo or spontaneous generation is most prominently manifested. Sofar, this seeming special potentiality of macerated hay has not been madethe subject of rigid and exhaustive inquiry, while the evidence recordedby Professor Tyndall concerning the extraordinary heat-resisting and infectiveproperties of hay-derived germs in their concrete form, is no doubtaccepted by the heterogenists as simply testifying to the possession bythis material of such potentiality. If, however, the disciples of heterogenyflatter themselves that from this last uninvaded vantage-ground they mayperadventure be able to withstand and bring discomfiture upon the advancinghosts of their opponents, the panspermists, they are destined tosignal disappointment.Within the last two years the animalcules produced so abundantly inhay-infusions have been the object of the author's special investigation,of this volumemany of the new species described in the systematic portionbeing, indeed, the outcome of such research.Hitherto the infusorial typescommonly observed in hay-infusions have been of comparatively largesize, belonging to the higher order of the Ciliata, and pertaining to suchgenera as Paramecium, Colpoda, Cyclidium, Oxytricha, and Vorticella.These, however, represent numerically but an insignificant minority comparedwith the vast hosts of flagellate forms which abound in a hay-infusionduring the earlier days of its maceration. In such infusions, watched fromday to day, and produced from hay obtained from different localities, thenumber of types presenting themselves in tolerably regular sequence wasfound to be perfectly marvellous. Foremost among the generic groupsputting in their appearance must be mentioned that of Heteromita, frequentlyrepresented simultaneously by three or four species, and including moreespecially the Heteromita (Manas) lens of O. F. Muller, H. caudata, andH. gracilis. Other genera, such as Oikomonas, Dinomonas, Petalomonas,Rhabdomonas, Ampkimonas, Monas proper, Cryptomonas, Hexamita, andGymnodinium, contribute likewise an almost equally considerable contingentwhile Bacteria in their characteristic motile and;quiescent states areinvariably present, and furnish an abundant and ever ready pabulum fortheir more highly organized animal consociates.The question presented to the author for solution was, from whencewere derived all these myriad organisms, frequently produced in such
136 SPONTANEOUS GENERATION.abundance as to literally jostle each other for room in every drop of waterextracted for examination? The heterogenists, including notably MM.Pouchet and Pennetier, have asserted that the highest Ciliate types presentin hay-infusions, such as Colpoda and Vorticclla, are generated de novo outof the filmy pellicle, or so-called " proligerous membrane," that in the courseof a few days makes its appearance upon the surface of the liquid. Thisproligerous membrane, again, is further represented to be formed from theaccumulated dead and floating bodies of Monads, Bacteria, and Vibrios,that first appeared in the infusion, and to constitute a kind of primordialstroma or pseud-ovary, out of the granular constituents of which, throughcoalescence at various points of the component particles, true eggs aredeveloped, giving birth to such Ciliata as sooner or later appear uponthe scene. The Monads themselves are treated as primary motile molecules,occupying a place midway between the organic and inorganic, andpossessing motile properties most nearly corresponding with the molecularor Brownian motions of minute inorganic particles. The spontaneousderivation of these Monads from the dead and disintegrated particles ofthe macerated hay is regarded as too obvious to need discussion. It hasbeen positively ascertained by the author, however, that these minutebeings are derived from spores which literally encrust with their countlessnumbers the stalks and blades of the vegetable matter ;these again beingthe product of pre-existing monad forms, whose active life was passed inclose association with the green and growing hay under the circumstanceshereafter narrated.In order to arrive at a comprehensive insight into the life-phenomenaand progressive developmental manifestations of the specialgroup of infusorialanimalcules now under consideration, hay from different localitieswas placed in maceration and examined continuously, from its first contactwith the fluid medium, for periods varying in duration from a few days onlyto several weeks. The water added to the hay was of the purest possibledescription, and was frequently boiled for some time to prevent the introductionof extraneous germs. In all instances, the results obtained werebroadly and fundamentally the same, and differed only with respect tothe specific types found living together in the separate infusions. Evenhere, however, the general dominance of two or more special forms wasnotably apparent. Commencing with the first wetting and simultaneousexamination of any given sample, spores of different sizes were foundcongregated in countless numbers, and in various orders of distribution,throughout the surfaces of the vegetable tissues. The majority of thesespores were excessively minute, spherical, of the average diameter of thel-2O,oooth part of an English inch, and required necessarily the employmentof the highest powers of the compound microscope for the correct registrationof their characteristic form and size.Sometimes these spores were tobe observed collected in definite spherical heaps, but more often they werescattered in irregular-shaped patches, such patches being often again more
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aoamoa
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"Our little systems have their day,
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TOTHOMAS HENRY HUXLEY, LL.D.,F.R.S.
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viiiPREFACE.experience some disappo
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XPREFACE.ready and valuable assista
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LEEUWENHOEtfS OBSERVATIONS. 3relate
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LEEUWENHOEICS OBSERVATIONS.5spatter
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LEEUWENHOEK'S OBSERVATIONS.Jstopped
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SIfi E. KING, 1693. JOHN HARRIS, 16
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STEPHEN GRA Y, 1696. LEEUWENHOEK, 1
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HENRY BAKER, 1742, 1753.13"Oct. 6th
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O. F. MULLER, 1773-1786. 15ledge of
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EHRENBERG, 1836. 17Notwithstanding
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F. DUJARDIN, 1841. T. VON SIEBOLD,
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FRIEDRICH STEIN, 1849-1854. 21cules
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CLAPAREDE AND LACHMANN, 1858-1860.
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F. STEIN, 1859. R. M. DIES ING, 184
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ANDREW PRITCHARD, 1861. H. JAMES-CL
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DALLINGER AND DRYSDALE, 1873-1875.
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CHAPTER II.THE SUB-KINGDOM PROTOZOA
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AFFINITIES OF THE SPONGIDA. 33ordin
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PRIMARY SUBDIVISIONS A UTHOKS S YST
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AUTHORS PHYLOGENETIC SCHEME. 37DIAG
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FLA CELLA TA -PANTOS TOMA TA ; FLA
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CHOANO-FLAGELLATA; MYCETOZOA. 41acc
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MYCETOZOAj LABYRINTHULIDA. 43From t
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GROUPS PROTISTA AND MONERA. 45of th
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DISTINCTION BETWEEN PROTOZOA AND PR
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( 49 )CHAPTER III.NATURE AND ORGANI
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AUTHORS CLASSIFICATORY TABLE.TABULA
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UNICELL ULAR NA TURE. 5 3dissolutio
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UN1CELL ULAR NA TURE. 5 5of the ent
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CUTICULAR ELEMENTS. 57substance the
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EXCRETED ELEMENTS. 59by the interca
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EXCRETED ELEMENTS. 6 1transparent,
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ENCYSTMENT. 63corresponding type of
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LOCOMOTIVE AND PREHENSILE APPENDAGE
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ORAL APERTURE. 67Oral Aperture or C
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CONTRACTILE VESICLES. 69shadowed. A
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CONTRACTILE VESICLES.71in the major
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NUCLEUS OR ENDOPLAST. 73to indicate
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NUCLEUS OR ENDOPLAST. 75Spirostomit
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NUCLEOLUS OR ENDOPLASTULE. 77with t
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COLOURING SUBSTANCES. 79held to ind
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TRICHOCYSTS. 8 1crowded together an
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TRICHOCYSTS. 83follow it,and being
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NATURE AND AFFINITIES OF THE SPONGE
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( 195 )CHAPTER VI.SYSTEMS OF CLASSI
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CLASSIFICATION OF THE INFUSORIA.197
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MULLERS CLA SSIPICA TOR Y S YSTEM.
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EHRENBERG'S CLASSIFICATORY SYSTEM.2
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CLASSIFICATORY SYSTEMS OF SIEBOLD A
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CLAPAREDE AND LACHMANWS CLASSIFICAT
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DIESINGS CLASSIFICATORY SYSTEM. 207
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S TEIN'S CLA SSIFICA TOR Y S Ki TEM
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A UTHOKS CLASSIFICA TOR Y S YSTEM.
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A UTHOR'S CLA SSIPICA TOR Y S YSTEM
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A UTHOKS CLASSIPICA TOR Y S YSTEM.
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CLASS FLAGELLA TA. 2 I7more extensi
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GENUS TRYPANOSOMA. 219Trypanosoma s
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GENUS MASTIGAMCEBA . 221The some ha
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;,HAB.GENUS REPTOMONAS. 22$immediat
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ORDER RA DIO-FLA CELLA TA.225Podost
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Body subspherical orGENUS ACTINOMON
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GENUS SPONGASTERISCUS. 229Spongocyc
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Cladomonas.ipidodendrtSpongomonas.D