GENETIC REPRODUCTION. 97be essentially hermaphrodite. Conjugation under any circumstance is,however, interpreted by Engelmann as productive of a mere revitalizingor rejuvenating power, enabling the unicellular organism to continue itsnormal mode of multiplication by binary division. During all conjugativeprocesses, the disintegration and subsequent reconstruction of the endoplasticelement, and the assumption by the endoplastule of a striatedaspect, reported by Biitschli, is entirely corroborated.While inclined to acquiesce with the views of Engelmann and Biitschlinow briefly recorded, so far as their interpretation of the conjugative processis concerned, and, as previously intimated, to attribute to this processa chiefly revitalizing or rejuvenating function as implying the most logicalpossible interpretation of the reproductive phenomena exhibited by unicellularorganizations, the present author is not prepared to follow theseauthorities in that part of their argument which involves the disassociationof the endoplast with any function beyond that possessed by the nucleus ofan ordinary tissue-cell, or to deny to the Infusoria any faculty of reproductionbeyond that of binary division. The one alternative is mostintimately, and indeed inseparably, involved with the other, and in thisconnection sufficient consideration has certainly not been granted byBiitschli and Engelmann to the results practically, and not merelytheoretically, obtained by other workers in the same field. That theendoplast does, under certain conditions, exhibit phenomena entirelydistinct from those presented by the simple histologic nucleus, and thatthe Infusorial organism can propagate its kind by means other than thoseof simple fission, has been already demonstrated in the section devoted toexternal and internal gemmation, and is conspicuously manifested in suchtypes as Hemiophrya gemmipara among the Acinetidse, in which ramifyingdiverticula of the endoplast ascend into the anterior bud-like extensions ofthe body-substance and become separated off, and enclosed within the same,when the latter are severed from the parent organism ; phenomena ofan essentially similar kind are likewise presented by the gemmiparousAcineta mystacina and various species of the genus Spirochona. Thereis, further, no sufficient reason for doubting the accuracy of the observationsfirst made by Eckhard in the year 1846, and since confirmed andextended by Claparede and Lachmann, concerning the production ofinternal embryos in Stentor cceruleus and polymorphic, these embryos beingdeveloped individually from a single element or node of the characteristicmoniliform endoplast. Where such embryos have been reported, Engelmannand Biitschli have dismissed them as merely externally derivedparasitic forms, mostly Acinetae, referable to the genus Spkaropkrya ofClaparede and Lachmann, synonymous with Engelmann's genus Endosph(zra.Such in many cases they undoubtedly are, and notably in thosesupposed embryonic forms associated by Stein with the several typesStentor polymorphic, Bursaria truncatella, and Stylonychia mytilus /_. theseunmistakable parasitic forms represent, however, but a small and altogetherII
98 ORGANIZA TION OF THE INFUSORIA.unimportant minority. Undoubted evidence of the production of embryosthrough the breaking up of the substance of the endoplast is afforded, inaddition to the instances already cited, by Haeckel, in his account andillustrations of the pelagic form Codonella campanella, and by Balbiani, inconnection with his most recent investigations of the Miillerian typeDidinium nasutum. In the genus Euglena and its allies, the production ofembryos through the increment in size and splitting up of the substance ofthe endoplast has been amply demonstrated by Stein and Carter, andis confirmed also by the observations of the present author. As a lastillustration of this production of germs or embryos through the subdivisionof the endoplastic element, may be cited Professor Allman's account ofthe development of a species of Epistylis, communicated to the meeting ofthe British Association, held at Brighton in the year 1875. In this caseindividuals possessing a normal development of the ordinary band-likeendoplast were observed to undergo encystment, the enclosing membranebecoming after the lapse of a few days so opaque as to preclude a clearview of its contents. Upon breaking the capsule open, however, it wasfound that the endoplast had increased prodigiously at the expense ofthe surrounding plasma, presenting the aspect of a long and much convolutedcord, while in still more advanced phases this cord-like endoplasthad separated into a number of ciliated and free-swimming germs whosecontour, as in the case of Didinium nasutum, most closely resembled theadult form of Trichodina (Halteria) grandinella. Reproductive phenomena,closely corresponding with those just described, have been recently reportedby Everts of Vorticella nebulifera.Before finally dismissing the subject of the production of embryos fromthe substance of the endoplast, a protest must undoubtedly be lodgedagainst Biitschli's assertion that the fragments into which it becomesseparated during or after conjugation, are merely cast out of the body aswaste matter. Such an unprofitable destinyis not usually found associatedwith so essential a structure, and in place of the purely negative evidenceadduced, it is equally probable, and far more logical, to presume thatthese detached fragments represent germs which ultimately develop tothe parent forms, as in the case of the endoplastic fragments of Stentor,Vorticella, Euglena, and other types already alluded to. Whether ornot the conjugation or fusion of two individual animalcules exerts a directinfluence upon the sporular or gemmiparous reproductive phases, in additionto the more ordinary binary segmental one, is as yet scarcely determ inable,though that in the first-named case it commonly, if not more usually,precedes spore-production is made evident through the investigations ofMessrs. Dallinger and Drysdale, added to those of the present author, inconnection with a considerable number of Flagellata.With respect to the actual act of conjugation, fusion, or zygosis, as itvariously denominated, it is worthy of remark that where such conjugationiscomplete and permanent, as in the family of the Vorticellidae, and so faris
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aoamoa
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"Our little systems have their day,
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TOTHOMAS HENRY HUXLEY, LL.D.,F.R.S.
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viiiPREFACE.experience some disappo
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XPREFACE.ready and valuable assista
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LEEUWENHOEtfS OBSERVATIONS. 3relate
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LEEUWENHOEICS OBSERVATIONS.5spatter
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LEEUWENHOEK'S OBSERVATIONS.Jstopped
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SIfi E. KING, 1693. JOHN HARRIS, 16
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STEPHEN GRA Y, 1696. LEEUWENHOEK, 1
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HENRY BAKER, 1742, 1753.13"Oct. 6th
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O. F. MULLER, 1773-1786. 15ledge of
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EHRENBERG, 1836. 17Notwithstanding
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F. DUJARDIN, 1841. T. VON SIEBOLD,
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FRIEDRICH STEIN, 1849-1854. 21cules
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CLAPAREDE AND LACHMANN, 1858-1860.
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F. STEIN, 1859. R. M. DIES ING, 184
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ANDREW PRITCHARD, 1861. H. JAMES-CL
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DALLINGER AND DRYSDALE, 1873-1875.
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CHAPTER II.THE SUB-KINGDOM PROTOZOA
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AFFINITIES OF THE SPONGIDA. 33ordin
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PRIMARY SUBDIVISIONS A UTHOKS S YST
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AUTHORS PHYLOGENETIC SCHEME. 37DIAG
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FLA CELLA TA -PANTOS TOMA TA ; FLA
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CHOANO-FLAGELLATA; MYCETOZOA. 41acc
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MYCETOZOAj LABYRINTHULIDA. 43From t
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GROUPS PROTISTA AND MONERA. 45of th
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( 195 )CHAPTER VI.SYSTEMS OF CLASSI
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CLASSIFICATION OF THE INFUSORIA.197
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MULLERS CLA SSIPICA TOR Y S YSTEM.
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EHRENBERG'S CLASSIFICATORY SYSTEM.2
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CLASSIFICATORY SYSTEMS OF SIEBOLD A
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S TEIN'S CLA SSIFICA TOR Y S Ki TEM
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A UTHOKS CLASSIFICA TOR Y S YSTEM.
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CLASS FLAGELLA TA. 2 I7more extensi
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GENUS TRYPANOSOMA. 219Trypanosoma s
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;,HAB.GENUS REPTOMONAS. 22$immediat
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ORDER RA DIO-FLA CELLA TA.225Podost
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Body subspherical orGENUS ACTINOMON
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GENUS SPONGASTERISCUS. 229Spongocyc
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Cladomonas.ipidodendrtSpongomonas.D