ENCYSTMENT. 63corresponding type of form with certain modifications, the areas betweenthe longitudinal ridges being transversely striate or otherwise ornamented,is common to the encystments of various Vorticellidae, while a somewhatsimilar one, having the elevated crests transversely placed, recurs in theAcinete type Podophrya fixa. In many species, such as Stylonychia pustulataand Pleurotricha lanceolata, the exterior surface of the spheroidal cystis closely studded with more or less irregular papilliform elevations. Inall these instances of simple or protective encystment, the animalcule uponthe return of favourable conditions assumes once more its normal aspect,and, breaking through the walls of its temporary prison-house, resumesits customary active habits. It is undoubtedly to the possession of thissimple self-protective faculty that the extensive distribution and prolongedvitality of many infusorial forms is mainly due. Where the ponds, ditches,or other tracts of fluid containing such animalcules become entirely dried up,these latter run no risk of extermination. Throwing out around them theirtransparent envelopes, they remain in a quiescent or torpid state until thereappearance of the previous congenial surroundings, or, taken up by thepassing breeze, are wafted away in the form of dust until conditions areencountered corresponding sufficiently with those under which they originallyflourished.The encystment of the second order to be mentioned is of comparativelyrare occurrence, and, instead of being associated with a simple conservativefunction, is an accompaniment of, or rather the prelude to, the phenomenonof multiplication by binary division, and may for this reason be mostappropriately denominated " duplicative encystment." The preliminarymanifestations and aspect of the constructed cyst correspond essentiallywith those recorded of the simply protective form but the; animalculeenclosed within its capsule, instead of resolving itself into a quiescent andinert mass, divides itself by the ordinary mode of increase by transversefission, the two halves shortly after making their exit through the wallsof the cyst, smaller in size, but in all other respects corresponding structurallywith the single pre-existing zooid. Encystment of this specialtype, in conjunction with other noteworthy data, has been observed byClaparede and Lachmann of the Holotrichous form Amphileptus meleagris.As recorded in detail later on, this animalcule is essentially predatory inits habits, and is addicted to preying upon the stationary and defencelesszooids of the Vorticellidan genus Epistylis,in the same manner that theMyxopod Vampyrella feeds upon the frustules of the Diatom Gomphonema.In a like manner also, having gorgeditself to satiety, the devouringA mphilepttis builds its cyst on the apex of the supporting stem of its latestvictim, and there undergoes the metamorphosis above described. Duplicativeencystment is recorded by Stein of Glaucoma scintillans, and in accordancewith the observations of that authority occurs also in Colpoda cucullulus,in combination with that variety of the process next described.The third, and remaining form of encystment to be enumerated, closely
64 ORGANIZA TION OF THE INFUSORIA.approaches in many instances the one last mentioned, while in others itexhibits widely distinct features. Like the preceding,it is connected withthe phenomena of reproduction, but the encysted animalcule multipliesitself not merely by the process of binary fission, but by the subdivision ofthe encapsuled mass into a greater or less number of spore-like bodieswhich, after a more or less prolonged quiescent state, developto theparent form. This type of encystation may be most appropriatelydenominated "sporular encystment," and the cyst or capsule secretedin such instances, a " sporocyst." Details of this special mode of multiplicationare given in the section devoted to the subject of reproduction,and it isonly requisite here to indicate one important point in which suchsporular encystment departs widely from both of the preceding kinds.In each of these latter the cyst or capsule producedis the product of aprimarily single and independent animalcule, but in the one now alludedto itvery frequently, though not invariably happens, that such a cyst isthe product of two primarily amalgamated or conjugated zooids.manyIn certaincases, even, as, for example, Heteromita uncinata, as as three or fourconjugated animalcules build up the characteristic sporocyst. This specialsporular form of encystment is, with but few exceptions, limited to theFlagellate class of the Infusoria.Locomotive and Prehensile Appendages.All of the variously modified appendages possessed by the severalorders of the Infusoria, used indifferently for the purposes of locomotion orprehension, are to be regarded as mere extensions of the body-protoplasm ;sometimes, as in most Flagellata, they are produced directly from the externalsurface of the ectoplasm, and in others, as the Ciliata, from the deeperor cortical layer of that element. In certain Tentaculifera the characteristictentacle-like appendages would seem to originate in close proximity to thecentral or endoplasmic region. With the exception of the organs lastmentioned, which would appear to most nearly represent specialized modificationsof the pseudopodia of the Radiolaria, the transition from one tothe other of the several types of appendages borne is most distinct andgradual. In this manner, flagella can be characterized only as isolated andmore or less elongate cilia while the divers forms of;setse, styles, and uncinipossessed most abundantly by the Ciliate section of the series, can be regardedas variations only, in separate directions, of similar simple cilia. Viewedfrom an independent standpoint, and as is requisite for the purposes oftechnical diagnosis, the term of " cilia " may be conveniently restricted tosuch short, slender, vibratile appendages as constitute the ordinary locomotiveorgans of a Paramecium, or the more or less convolute adoral ciliarywreath of a Vorticella. With the name of "setae" are to be associated theslender, hair- like, more or less flexible but non-vibratile appendages thatclothe the entire body of a Pleuronema, that are developed girdle-wise, andfulfil a special leaping function in the genus Halteria, or that in an isolated
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aoamoa
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"Our little systems have their day,
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TOTHOMAS HENRY HUXLEY, LL.D.,F.R.S.
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viiiPREFACE.experience some disappo
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XPREFACE.ready and valuable assista
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LEEUWENHOEtfS OBSERVATIONS. 3relate
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LEEUWENHOEICS OBSERVATIONS.5spatter
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LEEUWENHOEK'S OBSERVATIONS.Jstopped
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SIfi E. KING, 1693. JOHN HARRIS, 16
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STEPHEN GRA Y, 1696. LEEUWENHOEK, 1
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- Page 28 and 29: O. F. MULLER, 1773-1786. 15ledge of
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- Page 34 and 35: FRIEDRICH STEIN, 1849-1854. 21cules
- Page 36 and 37: CLAPAREDE AND LACHMANN, 1858-1860.
- Page 38 and 39: F. STEIN, 1859. R. M. DIES ING, 184
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PRESER VA TION. I 13Preservation of
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ME THODS OF INVES TIGA TION. I I5fo
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METHODS OF INVESTIGA TION.1 1^only
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RED I; TUBERVILLE NEEDHAM. 119simil
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JOHN HARRIS ; SPALLANZANI.l 2 rfor
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LORENZ OKEN; EHRENBERG. 123moment w
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POUCHET ; PASTEUR. 125Director of t
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PROFESSOR TYNDALL. 127now to be sup
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PROFESSOR TYNDALL. I29needle-dip fr
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PROFESSOR TYNDALL. 131tubes, as giv
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DALLINGER AND DRYSDALE. 133four day
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A UTHOKS INVESTIGA TIONS. 13 5and D
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A UTHORS INVESTIGA TIONS. 13 7or le
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AUTHOR'S INVESTIGATIONS. 139and lai
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A UTHORS INVESTIGA TIONS. 1 4 1clos
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( 143 )CHAPTER V.NATURE AND AFFINIT
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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NATURE AND AFFINITIES OF THE SPONGE
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( 195 )CHAPTER VI.SYSTEMS OF CLASSI
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CLASSIFICATION OF THE INFUSORIA.197
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MULLERS CLA SSIPICA TOR Y S YSTEM.
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EHRENBERG'S CLASSIFICATORY SYSTEM.2
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CLASSIFICATORY SYSTEMS OF SIEBOLD A
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CLAPAREDE AND LACHMANWS CLASSIFICAT
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DIESINGS CLASSIFICATORY SYSTEM. 207
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S TEIN'S CLA SSIFICA TOR Y S Ki TEM
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A UTHOKS CLASSIFICA TOR Y S YSTEM.
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A UTHOR'S CLA SSIPICA TOR Y S YSTEM
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A UTHOKS CLASSIPICA TOR Y S YSTEM.
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CLASS FLAGELLA TA. 2 I7more extensi
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GENUS TRYPANOSOMA. 219Trypanosoma s
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GENUS MASTIGAMCEBA . 221The some ha
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;,HAB.GENUS REPTOMONAS. 22$immediat
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ORDER RA DIO-FLA CELLA TA.225Podost
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Body subspherical orGENUS ACTINOMON
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GENUS SPONGASTERISCUS. 229Spongocyc
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Cladomonas.ipidodendrtSpongomonas.D