Cecropiaceae: Coussapoa and Pourouma, with an ... - CNCFlora

Coussapoa
Type. Colombia. Antioquia: Jardin, 1800 m, Dec
1941 (2), Bro. Daniel 2652 (holotype, US; isotypes,
COL, F).
Coussapoa macarenensis Cuatrecasas, Caldasia 7: 292.
1956. Type. Colombia. Meta: Sierra de la Macarena,
Caino Entrada, 24 Dec 1949 (5), Phillipson & Idrobo
1914 (holotype, BM; isotypes, COL, US).
Coussapoa macarenensis Cuatrecasas var. antioquiensis
Cuatrecasas, Caldasia 7: 293. 1956. Type. Colombia,
Antioquia: Rio Cauca, Puento Valdiva, 17-
20 Feb 1942 (2), Metcalf& Cuatrecasas 30093 (holotype,
F; isotypes, A, MO, US).
Coussapoa mutisii Killip & Cuatrecasas, Caldasia 7:
293. 1956. Type. Colombia. Without locality, - lutinous or to (sub)hirsute, often also with reddish-brown
pluricellular hairs and sometimes
arachnoid hairs; perianth ca. 1 mm high, densely,
minutely puberulous; stamens two, (rather) far
exceeding the perianth. Pistillate inflorescence
mostly unbranched, sometimes poorly branched;
heads 1(-4), free or occasionally partly fused,
(sub)globose, ca. 5-10 mm, in fruit up to 40 mm
diam.; (common) peduncle (1-)2-13 cm long, 1-
4 mm thick, densely to sparsely puberulous to
hirtellous to shortly velutinous or to (sub)hirsute,
(2),
Mutis 646
often also with reddish-brown
(holotype, US).
pluricellular hairs
Coussapoa planitiensis Cuatrecasas, Caldasia 7: 295. and sometimes arachnoid hairs; perianth ca. 1-
1956. Type. Colombia. Vaupes: Rio Guayabero, 8 2 mm high, densely minutely white to brownish
Nov 1939 (6), Cuatrecasas 7510 (holotype, US). puberulous; fruiting perianth at the apex green,
below the apex orange. Interfloral bracts
(sub)spathulate to subpeltate, white to brownish
puberulous at the apex.
Distribution (Fig. 9). Central America (from
Guatemala to Panama), the Andean region (Ecuador
to Venezuela), extending to the coastal
mountain range of Venezuela, and wide-spread
(especially) in the (Upper) Amazon Basin; in upland,
riverine, and mountain forest, up to 2400
m; often in secondary vegetation.
Tree or shrub, hemi-epiphytic or terrestrial, up
to 35 m tall. Leafy twigs 5-15 mm thick, sparsely
to densely white (to brown) puberulous to hirtellous
to (sub)hirsute to (sub)villous. Lamina
coriaceous, broadly ovate to subovate, sometimes
to elliptic, occasionally to obovate, 6-60
x 5-40 cm, apex obtuse to acute, sometimes
rounded to emarginate, base obtuse to cordate
(sometimes deeply cordate with overlapping
lobes), margin entire to subcrenate; upper surface
glabrous, lower surface sparsely to densely minutely
puberulous in the areoles and mostly also
on the reticulum, hirtellous to hispidulous on the
(parallel) tertiary venation, sometimes puberu-
Specimens studied. GUATEMALA. IZABAL: Peten
road, 2 km from Fronteras, Rio Juan Vicente, 11 May
1971 (8), Contreras 10763 (DUKE, MO, P, US); Quirigua,
15-31 Mar 1922 (6), Standley 23766 (GH, NY),
lous to (sub)hirsute, on the midrib and the lateral 26-27 Apr 1939 (9), Standley 72288 (F, NY); Lago
veins sparsely, sometimes rather densely, puber- Izabal, between Punta dos Reales and Punta de Leulous
to hirtellous or to (sub)hirsute, the whole
chuga, 17 Apr 1940 (2), Steyermark 39618 (F).
HONDURAS. ATLANTIDA: La Ceiba, 22 Sep 1916
surface covered with sparse to dense whitish to (9), Dyer A85 (F, US); Lancetilla Valley, nr. Tela, 6
brownish subpersistent arachnoid hairs; lateral Dec 1927-20 Mar 1928 (st), Standley 53230 (A, F,
veins (7-)9-24, straight or slightly curved, main US), (9), Standley 53991 (A, F, US), (st), Standley 54325
basal pair branched, reaching the margin below (A, F), (2), Standley 55155 (A, F, US), (st), Standley
55445 (A, F,
the middle of the lamina, in
US), (6), Standley 56623
large leaves
(F). COMAYAGUA:
some- Lake Yojoa, Pito Solo, nr. Pito Solo, 16 Apr 1951 (6),
times also with other lateral veins poorly P. H. Alien 6186 (F, GH), 8 Aug 1932 (st), Edwards
branched (furcate); intercostal venation more or 400 (A, F, US); 15 km E of Lake Yojoa, 20 May 1974
less prominent; petiole (2-)3-20 cm long, white (6), Hazelett s.n. (MO); nr. Pito Solo, 14 Aug 1973 (2),
Hazelett 664
(to yellowish) puberulous to hirtellous to
(MO); Lake Yojoa, 29 Jul-10 Aug 1951
(6), Howard et al. 605 (A); nr. Pito Solo, 18 Apr 1945
(sub)hirsute or to (sub)villous; stipules 3-20 cm (2), J. V. Rodriguez 2929 (F); Rio Tempemechin, ca.
long, puberulous to hirtellous to strigose to hir- 6 km N of Pito Solo, 15 Apr 1951 (d), L. O. Williams
sute, mostly also with dense (moniliform) red- et al. 18014 (F, GH). COPAN: Rio Copan, 2 mi E of
dish-brown pluricellular hairs and sometimes Copan ruins, 27 Aug 1975 (8), Molina et al. 30747 (F,
MO). CORTES: Naciemento del Rio
arachnoid
Lindo, nr.
hairs. Staminate
Jaral, 9
inflorescence Apr 1947 (6), Standley et al. 7081 (F); Rio Londo, 8
branched, occasionally unbranched; heads (1-)2- Apr 1945 (6), L. O. Williams et al. 12394 (F, GH); Rio
30(-ca. 50), often partly fused, (sub)globose to Lindo, nr. Carrizal, 12 Apr 1951 (st), L. O. Williams
ovoid to oblate, often more or less irregular in et al. 17817 (F), GRACIAS A Dios: Rio Platano, 0-4
hours
shape and dimensions, ca. 5-10(-20) mm upstream Ras, 23 May 1973 (9), Gentry et al.
diam.; 7530 (MO, U).
common peduncle 2-6 cm long, densely to NICARAGUA. BLUEFIELDS: Finca Santa Rosa, ca.
sparsely puberulous to hirtellous to shortly ve- 2.5 km ENE of Rama, 5 Apr 1966 (6), Proctor et al.
101