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Cecropiaceae: Coussapoa and Pourouma, with an ... - CNCFlora

Cecropiaceae: Coussapoa and Pourouma, with an ... - CNCFlora

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90 Flora Neotropica<br />

<strong><strong>an</strong>d</strong> Mocari, 150 m, 6 Mar 1987 (6), Fonnegra et al. cm long, glabrous; peri<strong>an</strong>th 1-2 mm high, gla-<br />

1798 (BG, HUA). VICHADA: Gaviotas, Caino Ariba, 19<br />

Feb 1973 (9), Cabrera 2668 brous; fruiting peri<strong>an</strong>th yellow or or<strong>an</strong>ge. Inter-<br />

(COL).<br />

VENEZUELA. AMAZONAS: Rio Cucucucuma, be- floral bracts (sub)spathulate, sparsely puberulous<br />

low Raudal Pacure, 9 Nov 1950 (6), Maguire et al. at the apex.<br />

29408 (NY); Dpto. Atabapo, 15 km SE of S<strong>an</strong> Fer- Distribution (Fig. 8). Southern Mexico <strong><strong>an</strong>d</strong><br />

n<strong><strong>an</strong>d</strong>o de Atabapo, 10-16 Feb 1988 (d), Stergios et al. north-western<br />

11530<br />

Guatemala; in evergreen forest in<br />

(BG).<br />

BRAZIL. AMAZONAS: Mun. Sao Paulo de<br />

moist<br />

Olivenoa,<br />

places (often along streams) or in seminr.<br />

Palmares, 11 Sep-26 Oct 1936 (2), Krukoff 8273 deciduous forest; up to 1700 m.<br />

(A, F, G, K, LE, MO, NY, P, S, U, US, type collection<br />

of C. parvifolia), (2), Krukoff8539 (A, BM, F, G, LE, Specimens studied. MEXICO. CHIAPAS: Ov<strong><strong>an</strong>d</strong>o Mt.,<br />

MO, NY, P, S, U, US); Mun. Sao Paulo de 17 Dec 1936<br />

Olivenca,<br />

(6), Matuda 446 (MO, NY, US); S<strong>an</strong>ta<br />

Belem Creek, 26 Oct-11 Dec 1936 (6), Krukoff8658<br />

Rita, Mapastepec, J<strong>an</strong> 1938 (6), Matuda 2020 (A, F,<br />

(A, BM, F, G, K, LE, MO, NY, P, S, U, US), (6), Krukoff<br />

K, NA, NY); between Guatimoc <strong><strong>an</strong>d</strong> Cacahuat<strong>an</strong>, 3<br />

8897 Dec 1941<br />

(A, BM, F, G, K, LE, MO, NY, P, S, U, US, type<br />

(9), Mir<strong><strong>an</strong>d</strong>a 1737 (US); NW of S<strong>an</strong> Fercollection<br />

of C. cornifolia); rd. M<strong>an</strong>aus-Itacoatiara, km n<strong><strong>an</strong>d</strong>o, NW ofTuxtla, 1000 m, 2 Apr 1950 (9), Mir<strong><strong>an</strong>d</strong>a<br />

65-70, 23 Oct 1963 (8), Oliveira 2764 (IAN); rd. Ma-<br />

6164 (US); nr. El Ocote, 30 km NW of Ocozocu<strong>an</strong>tla,<br />

naus-Itacoatiara, km 61, Res. Flor. Walter Egler, 17 24 Mar 1951 (9), Mir<strong><strong>an</strong>d</strong>a 6273 (US), GUERERO: Distr.<br />

Dec 1968 (6), Pr<strong>an</strong>ce et al. 9054 Montes de<br />

(M, MO, NY, P, S,<br />

Oca, S<strong>an</strong> Antonio, 18 Apr 1938 (9), Hinton<br />

U); rd. M<strong>an</strong>aus-Porto Velho, km 510, 5 km N of Rio 14018 (K, NY, P, US). JALISCO: Sierra Madre Occi-<br />

Purusinho, 17 Oct 1974 (6), Pr<strong>an</strong>ce et al. 22999 (INPA,<br />

dental, NW of S<strong>an</strong> Sebasti<strong>an</strong>, Las Mesitas, Arroyo de<br />

NY, U, US). PARA: Rd. Cuiaba-S<strong>an</strong>tar6m, km las<br />

1417, Caaitas, 1700 m, 15 Mar 1927 (6), Mexia 1872 (A,<br />

Rio Itapacura, 25 Nov 1977 (6), Pr<strong>an</strong>ce et al. 25748 BM, F, GH, MO, NY, US). VERACRUZ: Paso del In-<br />

(MO, RB, U).<br />

genio, Jun 1971 (9), Calzada 325 (U); <strong>with</strong>out locality,<br />

1930 (st), Purpus s.n. (B, F, K); Zacualp<strong>an</strong>, Mar 1908<br />

This species belongs to a group <strong>with</strong> two sta- (6), Purpus 5996 (BM, F, GH, MO, NY, US), Mar 1930<br />

mens <strong><strong>an</strong>d</strong> leaf margins which mostly are revolute (8), Purpus 11161 (K), Apr 1928 (6), Purpus 11162 (F,<br />

at the base, comprising e.g., C.<br />

type collection), Jun 1930<br />

microcarpa <strong><strong>an</strong>d</strong><br />

(9), Purpus 11162 (A, K,<br />

MO), Sep 1930 (9), Purpus 11162 (C), Apr 1931 (9),<br />

C. microcephala.<br />

Purpus 11162 (C), May 1933 (6 <strong><strong>an</strong>d</strong> 9), Purpus 11162<br />

Pr<strong>an</strong>ce et al. 9054 is aberr<strong>an</strong>t in having (A), May 1934 (6 <strong><strong>an</strong>d</strong> 9), Purpus 11162 (K, US), Sep<br />

(sub)obovate leaves.<br />

1929 (8), Purpus 11182 (A).<br />

GUATEMALA. QUEZALTENANGO: Colomba, ca.<br />

1000 m, 28 Dec 1934 (6), Skutch 2023 (BM, F, GH,<br />

36. <strong>Coussapoa</strong> purpusii St<strong><strong>an</strong>d</strong>ley, Publ. Field NY). SAN MARCOS: Finca El Porvenir, on Potrero Ma-<br />

Mus. Bot. 8: 6. 1930. Type. Mexico. Veracruz: tas<strong>an</strong>, Rio Cabuis, Volc<strong>an</strong> Tajumulco, 1000-1300 m,<br />

Zacualp<strong>an</strong>, Apr 1928 (6), Purpus 11162 12 Mar<br />

(ho-<br />

1940 (9), Steyermark 37602 (F), 25 Mar 1943<br />

lotype, F). Fig. 48.<br />

(st), Steyermark 52334 (F).<br />

The species appears to be related to C. parviceps.<br />

Tree, hemi-epiphytic or terrestrial, often (?)<br />

deciduous, up to 20 m tall. Leafy twigs 3-6 mm<br />

thick, glabrous or puberulous on the scars of the<br />

stipules. Lamina chartaceous, ovate to elliptic or<br />

to obtuse, occasionally obovate, 4-16 x 1-7 cm,<br />

apex acute to acuminate to obtuse, base obtuse<br />

sometimes subacute or subcordate, margin en-<br />

tire; both surfaces glabrous; lateral veins 4-6 pairs,<br />

curved, basal pair unbr<strong>an</strong>ched reaching the mar-<br />

gin below, at, or slightly above the middle of the<br />

lamina; intercostal venation pl<strong>an</strong>e; petiole 1-5<br />

cm long, glabrous; stipules 1-3 cm long, glabrous<br />

or sparsely to densely appressed-puberulous.<br />

Staminate inflorescences br<strong>an</strong>ched; heads 2-6,<br />

globose, ca. 5-8 mm diam.; common peduncle<br />

2-4 cm long, glabrous; peri<strong>an</strong>th ca. 1 mm high,<br />

glabrous or sparsely minutely puberulous; sta-<br />

mens three, far exceeding the peri<strong>an</strong>th. Pistillate<br />

inflorescences unbr<strong>an</strong>ched; heads globose, 4-6<br />

mm, in fruit up to 10 mm diam.; peduncle 3-4<br />

37. <strong>Coussapoa</strong> scabra Akkerm<strong>an</strong>s & Berg, Proc.<br />

Kon. Ned. Akad. Wetensch. Ser. C 85(4): 460.<br />

1982. Type. Brazil. Rondonia: Rio Ouro Pre-<br />

to, affluent of Rio Pacaas Novos, 18 Sep 1923<br />

(9), Kuhlm<strong>an</strong>n 470 (HJBR 19836) (holotype,<br />

RB; isotypes, RB, U). Fig. 49.<br />

Tree. Leafy twigs 2-5 mm thick, puberulous<br />

to subhirsute; internodes solid. Lamina charta-<br />

ceous, elliptic to ovate or obovate, 4-10 x 2.5-<br />

7 cm, apex acute to shortly acuminate, base ob-<br />

tuse to acute, margin entire; upper surface, sca-<br />

brous, lower surface, scabridulous, sparsely to<br />

rather densely puberulous to hirtellous; lateral<br />

veins 6-13 pairs, straight to curved, basal pair<br />

unbr<strong>an</strong>ched, reaching the margin far below the<br />

middle of the lamina; intercostal venation slight-<br />

ly prominent; petiole 1-3 cm long, densely pu-

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