<strong>Coussapoa</strong> FIG. 28. <strong>Coussapoa</strong>floccosa: 1, leafy twig <strong>with</strong> pistillate inflorescences (Irwin 2058). latter in having a narrower lamina, a smaller number of lateral veins, <strong><strong>an</strong>d</strong> a relatively short petiole. Pistillate inflorescences are needed to decide whether this taxon is distinct from C. crassivenosa at the specific or only intraspecific level. 17. <strong>Coussapoa</strong> floccosa Akkerm<strong>an</strong>s & C. C. Berg, Proc. Kon. Ned. Akad. Wetensch. Ser. C, 85(4): 451. 1982. Type. Brazil. Minas Gerais: Vi9osa, 16 Nov 1935 (2), Kuhlm<strong>an</strong>n 2074 (holotype, RB; isotypes, RB, U, US). Fig. 28. Tree or shrub, (mostly?) hemi-epiphytic. Leafy twigs 10-15 mm thick, (rather) densely 57 puberulous to (sub)hirsute. Lamina coriaceous, elliptic to ovate or obovate, 10-25 x 6-17 cm, apex rounded to obtuse or to emarginate, base obtuse to rounded to cordate, margin entire; upper surface often densely covered <strong>with</strong> white arachnoid hairs which later disappear, lower surface hirtellous to subtomentellous on the veins, the whole surface sparsely to densely covered <strong>with</strong> white arachnoid hairs, later deciduous; lateral veins 7-10 pairs, (main) basal pair distinctly br<strong>an</strong>ched, reaching the margin at or above the middle of the lamina; intercostal venation very prominent; petiole 1.5-5 cm long, tomentose to densely hirtellous; stipules 1-2.5 cm long, white to brownish appressed-puberulous to strig(ill)ose
58 Flora Neotropica to subsericeous (to subhirsute), sometimes also mm high, glabrous. Interfloral bracts (sub)peltate, <strong>with</strong> sparse arachnoid hairs. Staminate inflores- nearly glabrous. cences br<strong>an</strong>ched; heads numerous, globose, ca. 4 Distribution (Fig. 7). Central America: Costa mm diam.; common peduncle 1.5-4 cm long, Rica (Puntarenas), Nicaragua (Bluefields <strong><strong>an</strong>d</strong> Rio puberulous; peri<strong>an</strong>th ca. 1 mm high, puberulous; S<strong>an</strong> Ju<strong>an</strong>), P<strong>an</strong>ama (S<strong>an</strong> Blas); in wet (riverside stamens two, exceeding the peri<strong>an</strong>th. Pistillate or swamp) forest. inflorescences br<strong>an</strong>ched; heads 2-3, partly fused, (sub)globose, 5-7 mm diam.; common peduncle Specimens studied. NICARAGUA. BLUEFIELDS: Rio 1-3 cm long, densely puberulous; peri<strong>an</strong>th 1-2 Kama, between Kama <strong><strong>an</strong>d</strong> C<strong>an</strong>io Valentin, 10 Mar 1966 (2), Proctor et al. 27080 (F, US, type collection). mm high, glabrous. Interfloral bracts sub- Rio SAN JUAN: 1 km NW of Rio S<strong>an</strong>ta Cruz, 22 Feb spathulate, puberulous at the apex. 1984 (2), Moreno 23629 (BG). Distribution (Fig. 7). Brazil (Minas Gerais, Vi- COSTA RICA. PUNTARENAS: Peninsula de Osa, 8 qosa) in km S of forest. Rinc6n, 28 Feb 1965 (9), Jimenez 3016 (F). PANAMA. SAN BLAS: El Ll<strong>an</strong>o-Carti rd., km 19.1, Specimens studied. BRAZIL. MINAS GERAIS: Est. 1 Feb 1985 (2), Nevers et al. 4804 (WIS), 3 Nov 1985 Exp. de Agua Limpa, Feb 1968 (2), Gomes et al. 2819 (st), Nevers et al. 6186 (BG). SAN BLAS-PANAMA: El (RB); Vi9osa, 10 Nov 1958 (2), Irwin 2058 (F, NY, Ll<strong>an</strong>o-Carti rd., 28 Aug 1982 (d), Hamilton et al. 1062 R, US), 31 Aug 1934 (st), Kuhlm<strong>an</strong>n 1832 (US), 1935 (BG). (st), Kuhlm<strong>an</strong>n 2073 (RB, US), 16 Nov 1935 (2), Kuhlm<strong>an</strong>n 2074 (RB, U, US, type collection), 31 Aug 1934 This species appears to be one of a group which (st), Kuhlm<strong>an</strong>n 2117 (RB), 1935 (9), Kuhlm<strong>an</strong>n 2207 also includes C. <strong>an</strong>gustifolia <strong><strong>an</strong>d</strong> C. trinervia <strong><strong>an</strong>d</strong> (GUA), 8 Oct 1976 (9), G. Rodrigues et al. 877 (RB); is characterized by shortly petiolate, more or less Capolivinba, nr. Rio Novo, Sep 1895 (6), Schwacke distinctly obovate leaves <strong>with</strong> few lateral veins, 11895 (B, P). <strong><strong>an</strong>d</strong> sparse indumentum. This species, known only from the surroundings of Vicosa, is very closely related to C. cur- 19. <strong>Coussapoa</strong> herthae Mildbraed, Notizbl. Bot. r<strong>an</strong>ii; but the two taxa are sufficiently different Gart. Berlin 14: 29. 1938. Type. Ecuador. Pifor them each to be regarded as species in their chincha: S<strong>an</strong> Carlos de los Colorados, 5 Oct own right. 1935 (d), Schultze-Rhonhof 953 (holotype, B). Fig. 30. 18. <strong>Coussapoa</strong> glaberrima W. Burger, Phytolo- gia 26: 422. 1973; Burger, Fieldi<strong>an</strong>a Bot. 40: 133, t. 22. 1977. Type. Nicaragua. Bluefields: Rio Kama, between Kama <strong><strong>an</strong>d</strong> C<strong>an</strong>io Valentin, 10 Mar 1966 (2), Proctor et al. 27080 (holo- type, F; isotype, US). Fig. 29. Tree, hemi-epiphytic or terrestrial, small. Leafy twigs 2-5 mm thick, glabrous. Lamina subcoriaceous, obovate to narrow elliptic, 2-14 x 1-6 cm, apex acute, base acute to obtuse, margin entire; both surfaces glabrous; lateral veins 2-3 pairs, slightly curved, basal pair indistinctly br<strong>an</strong>ched, reaching the margin far above the middle of the lamina; intercostal venation slightly prominent; petiole 0.5-2 cm long, glabrous; stipules 1.2 cm long, glabrous. Staminate inflorescences br<strong>an</strong>ched; heads ca. 10-12, ca. 2 mm diam.; common peduncle 1.5-3 cm long, gla- brous; peri<strong>an</strong>th ca. 1 mm high, glabrous; stamens two, just exceeding the peri<strong>an</strong>th. Pistillate inflorescences poorly br<strong>an</strong>ched; heads 2-3, globose, ca. 6 mm, in fruit up to 15 mm diam.; common peduncle 2-4 cm long, glabrous; peri<strong>an</strong>th ca. 1 Tree, mostly hemi-epiphytic, up to 25 m tall. Leafy twigs 5-13 mm thick densely, yellow to brown, strigose to hirsute to subvillous. Lamina coriaceous, elliptic to ovate to obovate or to ob- long, 16-32 x 10-20 cm, apex acuminate to acute, base obtuse to rounded or occasionally subcordate, margin entire or subcrenate; upper surface glabrous, sparsely to rather densely pub- erulous to hirtellous or on the main veins to subsericeous, sometimes <strong>with</strong> white arachnoid hairs which soon disappear; lateral veins (4-) 8-11 pairs, curved, basal pair distinctly to faintly br<strong>an</strong>ched, reaching the margin usually below, sometimes at or just above the middle of the lamina, sometimes also other lateral veins br<strong>an</strong>ched, a distinct submarginal vein in the lower part of the lamina; intercostal venation (often very) prominent; petiole (1-)3-9 cm long, puberulous to hirtellous (to subhirsute); stipules 2- 6 cm long, densely reddish- to yellowish-brown subsericeous to subhirsute to subvillous. Staminate inflorescences br<strong>an</strong>ched; heads m<strong>an</strong>y, globose, 2-3 mm in diam.; common peduncle
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Organization for Flora Neotropica C
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Copyright ? 1990 The New York Botan
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2 Flora Neotropica ABSTRACT Berg, C
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4 Flora Neotropica COUSSAPOA Engler
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Coussapoa 107 FXG. 56. Coussapoafil
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Coussapoa FIG. 57. Coussapoa tolime
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Morphology ent revision, especially
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Morphology; Phenology; Pollination
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Distribution and Ecology ruginea, P
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Pourouma 117 Key to the Species and
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Pourouma 119 35. Leafy twigs hirsut
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Pourouma 121 65. Arachnoid hairs co
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Pourouma 123 white-arachnoid indume
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Pourouma125 restal "La Paragua," Ri
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Pourouma127 LE, MO, NY, P, S, U, US
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Pourouma ~ I I MINISTERIO DE AGRICU
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Pourouma 131 '~~~~~~~~~~~~~~~~~~~~~
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Pourouma 133 3. Smaller veins (reti
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Pourouma 135 1953 (st), Steyermark
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Pourouma 137 of the species range t
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Pourouma 139 This subspecies resemb
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Pourouma 15520 (F, GH, NY), (9), Cu
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_' "( O^^Jgy -' i biccoolr P. bicol
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Pourouma 145 ,J, ? ^T \Sr~~u. I . .
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Pourouma 147 Rio Jurua, Marary, 8 O
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Pourouma (U), 17 Feb 1980 (6), Berg
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Pourouma ~~~~~~~~~~~~~~~~~~~~~~~~..
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Pourouma 153 FG R.: 17. 411... . ur
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Pourouma 155 long, yellow-velutinou
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.? P. cucura P. cuspidats GP. 7rorP
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Pourouma159 Pourouma triloba Klotzs
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Pourouma 161 between Rio Chicanan a
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Pourouma 163 FIG. FLORA 76 Prom mn
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Pourouma 165 m.m,--- * _ . 1 2'. A
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+ AJ ++ P O R O ULMA tomentosa. FIG
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Pourouma 169 Varadouro do Morcego,
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Pourouma qlq, _ j PIF P. t osantosa
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Pourouma 173 465 (P); Trois Sauts,
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Pourouma 175 I0711~~~~~~~~~~~~~~~~~
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Pourouma 177 Specimens examined. VE
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Pourouma 11 '.ciLnnr'.mon-lik 2 3 4
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Pourouma 181 the areoles; lateral v
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Pourouma 183 Ak~~~~~~~~~~~~?IIk i.
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Pourouma 185 I I STPE ,E Fn CL1 |SO
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Pourouma 187 . . . FIG. 91. Pouroum
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Pourouma 189 upper Rio Madre de Dio
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Pourouma 191 bi f In i FIG. 92. Pou
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Pourouma (BG), 9 Dec 1977 (d), Gent
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Literature Cited 195 and distributi
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Exsiccatae 197 e. subsp. chocoana (
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Exsiccatae Dombey, J., s.n. (1-42).
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Exsiccatae 201 Korning, J. et al.,
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Exsiccatae 20); 15764 (1-40); 15791
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Exsiccatae 205 Weberbauer, A., 3639
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Index of Scientific Names 207 conto