Cecropiaceae: Coussapoa and Pourouma, with an ... - CNCFlora

Pourouma 147
Rio Jurua, Marary, 8 Oct 1900 (c), Ule 5266 (HBG, 1951 (e), Garcia-Barriga 14002 (holotype, US; iso-
MG).
type, NY).
BOLIVIA. BENI: Prov. Moxos, S of Moxos, 18 Sep
1976 (9), Meneces et al. 338 (MG, MO); Rurrenabaque, Tree, up to 30 m tall. Leafy twigs 4-8 mm
Oct 1921 (9), Rusby 1599 (F, GH, NY, US). PANDO: thick, densely to sparsely yellow-hirsute, these
Prov. Nicolas Suarez, Mukden, Jun-Dec 1979 (2), Izawa
21 long hairs often intermixed with much shorter,
(U).
whitish hairs, sometimes also with sparse, brown,
Vernacular names. Colombia. bochoa tsaha pluricellular hairs, sometimes (sub)glabrous or
(Kofan), caimaron, caimaron silvestre, uva, uva subvelutinous. Lamina entire, ovate to subovate
del monte, uvilla, uvo; Amazonas: gurucana. or elliptic to oblong and 6-20 x 2.5-12 cm or
Venezuela: cocura, sadajii. Peru. sacha uvilla, 3-lobed to -parted (or in subjuvenile material up
uvilla, suia; Pasco: shewantaqui. Brazil: cucura, to 5-parted, sometimes with the segments almost
cucuva, imbauba (or ambauva) mansa, imbafba petiolulate) and up to ca. 25 x 25 cm, coriaceous
(or ambauva) do vinho, mapati, puruma, puru- to subcoriaceous, apex acuminate, acute, apicma-y,
sucuuiba, uva. Bolivia. uva silvestre; Beni: ulate, or sometimes emarginate, base subcordate
tanaribe, uva del monte.
to truncate to rounded to subacute; upper surface
This species shows strong similarities to P. bi- ? scabrous, occasionally smooth, yellow-hirsute
color, especially to the subspecies digitata and to -hirtellous on the whole surface or only on the
scobina. It mainly differs in the greater number main veins, lower surface yellow-hirsute (to -hirof
segments of the leaves of adult specimens (nor- tellous) to -substrigose on the main veins, white
mally 7-11 versus normally 5-7 in subsp. digi- arachnoid hairs in the areoles and on the smaller
tata and subsp. scobina), the smooth upper sur- veins; lateral veins 10-20 pairs, the basal pair
face, and the sparser indument on the leafy twigs. branched, tertiary venation prominent beneath;
The lamina of subsp. digitata and subsp. scobina petiole 3-17 cm long, yellow-hirsute, intermixed
is sometimes smooth and subjuvenile specimens with short, whitish hairs; stipules 2-12.5 cm long,
may have up to 9 leaf segments, which makes outside yellow-hirsute to -strigose, inside gladistinction
of the taxa on the basis of morpho- brous, caducous. Staminate inflorescences up to
logical characters sometimes impossible. For- 12 cm long and 4.5 cm wide; peduncle 3-6.5 cm
tunately, the taxa are allopatric (or almost so, see long, peduncle and branches yellow-hirsute;
p. 139).
flowers sessile, rarely pedicellate, clustered, but
The lectotype ofP. cecropiifolia is selected from not in globose heads; tepals almost free, (sparsenine
Martius syntype collections listed above, ly) puberulent; filaments free, shorter than the
that of P. sapida from two Karsten (syntype) col- tepals. Pistillate inflorescences in fruit up to 19
lections.
cm long and 10.5 cm wide; peduncle 3-9 cm long,
The fully mature fruiting perianth of most (if peduncle and branches yellow-hirsute;flowers 20not
all) Pourouma species is suitable for human 25; pedicel up to 0.6 cm long, in fruit up to 1
consumption, but P. cecropiifolia is the only cm; perianth 0.3-0.9 mm long, hispidulous (or
species in cultivation (see p. 116).
hirtellous); stigma peltate, ca. 2 mm in diam.,
The fact that the species is so often found in whitish puberulous. Fruiting perianth blue-black,
(formerly) inhabited places suggests that the ellipsoid to oblongoid to subglobose, ca. 1-1.5
present distribution is (at least) partly anthro- cm long, hispidulous (or hirtellous).
pogenous.
Distribution (Fig. 74). Colombia (Amazonas
and Vaupes), Venezuela (Amazonas and Boli-
5. Pourouma cucura Standley & Cuatrecasas in var), Ecuador (Napo and Morona-Santiago), Peru
Cuatrecasas, Fieldiana Bot. 28(1): 211. 1951. (Loreto and Madre de Dios) and Brazil (Amapa,
Type. Venezuela. Amazonas: Capihuara, Alto Amazonas, and Mato Grosso), extending to east-
Casiquiare, 5 Jun 1942 (Q), LI. Williams 15812 ern Venezuela (Bolivar); in non-inundated or in
(holotype, F; isotypes, G, NY, RB, US). periodically inundated (varzea or igap6) forest;
Fig. 70. mostly at low altitudes, sometimes (in Ecuador)
up to ca. 1500 m.
Pourouma garciana Cuatrecasas, Caldasia 7:298. 1956.
Type. Colombia. Amazonas-Vaupes: Rio Apaporis, Specimens examined. COLOMBIA. AMAZONAS: Rio
between Rio Kananari and Rio Pacoa, 1-15 Dec Apaporis, above mouth of Rio Kananari, Soratama,