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Cecropiaceae: Coussapoa and Pourouma, with an ... - CNCFlora

Cecropiaceae: Coussapoa and Pourouma, with an ... - CNCFlora

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190 Flora Neotropica<br />

<strong>Pourouma</strong> napoensis <strong><strong>an</strong>d</strong> P. herrerensis have<br />

been included in this revision after submission<br />

of the (pre-review) m<strong>an</strong>uscript <strong><strong>an</strong>d</strong> are, therefore,<br />

placed at the end of the systematic treatment,<br />

<strong><strong>an</strong>d</strong> not after P. hirsutipetiolata <strong><strong>an</strong>d</strong> P. mollis,<br />

respectively.<br />

24. <strong>Pourouma</strong> napoensis C. C. Berg, Brittonia<br />

42: 59-65. 1990. Type. Ecuador. Napo: Re-<br />

serva Biologia Jatun Sacha, 14 Aug 1987 (6),<br />

Palacios 1874 (holotype, QCNE; isotypes,<br />

AAU, BG, MO, NY, QAME). Fig. 92.<br />

Tree, up to 25 m tall. Leafy twigs 1-2.5 cm<br />

thick, yellow-hirsute, pluricellular hairs absent.<br />

Lamina 7-fid to -parted, ca. 25-45 x 25-45 cm,<br />

coriaceous, apex acuminate, base deeply cordate<br />

<strong>with</strong> overlapping lobes; upper surface smooth to<br />

slightly scabrous, yellow-hirsute to -hirtellous on<br />

the whole surface or only on the main veins,<br />

lower surface yellow-hirsute to -hirtellous on the<br />

veins, arachnoid hairs in the areoles <strong><strong>an</strong>d</strong> on the<br />

reticulum; lateral veins 16-25 pairs; petiole ca.<br />

25-40 cm long, yellow-hirsute; stipules 10-16 cm<br />

long, outside yellow-hirsute, inside glabrous, caducous.<br />

Staminate inflorescences ca. 12-22 cm<br />

long, ca. 8-14 cm wide; peduncle 4-8 cm long,<br />

peduncle <strong><strong>an</strong>d</strong> br<strong>an</strong>ches yellow-hirtellous to -pu-<br />

In the shape, dimensions, <strong><strong>an</strong>d</strong> indument of the<br />

lamina this species resembles P. oraria. It differs<br />

from the latter in the staminate flowers <strong>with</strong> their<br />

connate tepals <strong><strong>an</strong>d</strong> fully connate filaments. These<br />

features suggest a close relationship to P. hirsu-<br />

tipetiolata.<br />

25. <strong>Pourouma</strong> herrerensis C. C. Berg, C<strong><strong>an</strong>d</strong>ollea<br />

44(2): 513. 1989. Type, Peru, Loreto, Prov.<br />

Requena, Jenaro Herrera, Reserva Forestal,<br />

tree 4/122, 2 Oct 1985 (d), Spichiger et al. 1995<br />

(holotype, G; isotype, BG). Fig. 93.<br />

Tree, up to 25 m tall. Leafy twigs 3-6 mm<br />

thick, minutely puberulous to hispidulous (<strong><strong>an</strong>d</strong><br />

+ scabrous) or also <strong>with</strong> much longer, yellow<br />

hairs, pluricellular hairs dark brown <strong><strong>an</strong>d</strong> rather<br />

sparse. Lamina entire, oblong to elliptic to ovate,<br />

8-20 x 3-11.5 cm, (sub)coriaceous, apex shortacuminate,<br />

base acute to rounded; upper surface<br />

sparsely appressed-puberulous on the midrib or<br />

the whole surface covered <strong>with</strong> long, yellow hairs,<br />

lower surface appressed-puberulous on the main<br />

veins or also <strong>with</strong> long yellow hairs, arachnoid<br />

hairs in the areoles <strong><strong>an</strong>d</strong> on the reticulum; lateral<br />

veins 9-14 pairs, basal pair unbr<strong>an</strong>ched or<br />

br<strong>an</strong>ched, the basal part of these veins forming<br />

the basal part of the leaf margin (thus not separated<br />

from the margin by mesophyll), tertiary<br />

venation slightly prominent; petiole 3-10 cm<br />

berulous; flowers (sub)sessile, in<br />

long, minutely puberulous or also <strong>with</strong> long, yelglobose<br />

heads, low<br />

these 7-12 mm diam.; peri<strong>an</strong>th, tubular, 1-2 mm<br />

hairs; stipules 3.5-8 cm long, caducous, outside<br />

high, puberulous at the apex; stamens 3-4, filadensely<br />

puberulous <strong><strong>an</strong>d</strong> <strong>with</strong> brown, pluricellular<br />

ments completley connate, 4-6 mm long. Pistilhairs,<br />

or also <strong>with</strong> long, yellow hairs,<br />

sometimes<br />

late inflorescences up to 10 cm long <strong><strong>an</strong>d</strong> up to<br />

only a few appressed ones, inside<br />

12 cm wide, br<strong>an</strong>ched; peduncle 3-5 cm sparsely short-pubescent. Staminate infloreslong,<br />

cences<br />

peduncle <strong><strong>an</strong>d</strong> br<strong>an</strong>ches yellow-hirsute to -hirtel- up to 10 cm long <strong><strong>an</strong>d</strong> up to 9 cm wide;<br />

lous; flowers ca. 40-70; pedicel 0.3-0.5 cm, in<br />

peduncle 2-6 cm long, peduncle <strong><strong>an</strong>d</strong> br<strong>an</strong>ches<br />

fruit up to 1.2 cm long; peri<strong>an</strong>th ca. 5 mm densely puberulous to short-velutinous or also<br />

long, <strong>with</strong><br />

yellow-hirsute to -hirtellous, sparsely so in the long, yellow hairs; flowers sessile or pedicellate,<br />

most of them in globose heads, numerous<br />

uppermost part; stigma peltate.<br />

<strong><strong>an</strong>d</strong> 3-4 mm diam. or less<br />

Distribution (Fig. 81). Ecuador (Napo), in non-<br />

(5-12) <strong><strong>an</strong>d</strong> 5-8 mm<br />

inundated forest, at low altitudes.<br />

diam.; tepals ca. 1 mm long, connate, forming a<br />

+ urceolate peri<strong>an</strong>th, densely hairy; filaments<br />

Specimens examined. ECUADOR. NAPO: Reserva exceeding the tepals, free. Pistillate inflorescences<br />

Biologica Jatun Sacha, Rio Napo, 8 km below Puerto up to 14 cm long <strong><strong>an</strong>d</strong> 9 cm wide; peduncle <strong><strong>an</strong>d</strong><br />

Misahualli, 4 Sep 1987 (2), Cer6n M. 2019 (BG); 8 km br<strong>an</strong>ches short-velutinous; flowers 7-25, in 1-4<br />

SE of Tena, 30 Aug 1960 (2), Grubb et al. 1545 (K,<br />

NY); Reserva Biologica Jatun Sacha, Rio Napo, 8 km<br />

+ distinct clusters, pedicels up to 0.7 cm long;<br />

below Puerto Misahualli, 2 Oct 1986 (2), Palacios 1337 peri<strong>an</strong>th yellow-velutinous, except for the apex;<br />

(BG), 14 Aug 1987 (a), Palacios 1874 (BG, type col- stigma subpeltate.<br />

lection of P. napoensis); Lumbaqui, 12 May 1987 (a), Distribution (Fig. 81). Peru (Loreto), in non-<br />

Pennington et al. 12244 (BG, K).<br />

inundated forest.<br />

Specimens examined. PERU. LORETO: Prov. Requena,<br />

Jenaro Herrera, Aug-Sep 1976 (6), Bernardi s. n.

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