<strong>Pourouma</strong> 165 m.m,--- * _ . 1 2'. A . .. . . ; FIG. 78. <strong>Pourouma</strong> hirsutipetiolata subsp. hispida. Leafy twig <strong>with</strong> staminate inflorescences (Little e a. 21225). on the midrib, in scabrous leaves hispidulous over the whole surface, lower surface rather sparsely appressed-puberulous to strigose on the (main) veins, arachnoid hairs in the areoles, usually extending to the midrib, but on the main veins ? disappearing <strong>with</strong> age; lateral veins 7- 25 pairs, the basal pair br<strong>an</strong>ched or sometimes (in elliptic leaves) unbr<strong>an</strong>ched, the basal part of these veins forming the basal part of the leaf margin (thus not separated from the margin by mesophyll), tertiary (<strong><strong>an</strong>d</strong> quarternary) veins often rather prominent beneath; petiole 2-26 cm long, white-puberulous or yellow-(sub)hirsute, <strong><strong>an</strong>d</strong> usually also white, arachnoid hairs; stipules (2-)6- 18 cm long, outside whitish-strigose to yellowhirsute or -hirtellous, <strong><strong>an</strong>d</strong> <strong>with</strong> dense to sparse,
166 Flora Neotropica white, arachnoid hairs, usually also pluricellular This species is closely related to P. mollis <strong><strong>an</strong>d</strong> hairs, inside glabrous or <strong>with</strong> dense or sparse, P. melinonii. It is distinguishable by the occuryellow(ish) hairs, caducous. Staminate inflores- rence of ? dense, arachnoid hairs on the leafy cences up to 12 cm long <strong><strong>an</strong>d</strong> 12 cm wide; pe- twigs, stipules, petioles, main veins at the lower duncle 2-5 cm long, peduncle <strong><strong>an</strong>d</strong> br<strong>an</strong>ches + leaf surface <strong><strong>an</strong>d</strong>/or the inflorescences, <strong><strong>an</strong>d</strong>, moredensely puberulous, often also <strong>with</strong> white, arach- over, by the lack or scarcity of pluricellular hairs noid hairs; flowers sessile or pedicellate, most of on the leafy twigs. them in (sub)globose heads, these 3-5 mm diam.; Five subspecies, partly allopatric, c<strong>an</strong> be rectepals 0.5-1 mm long, connate, forming a more ognized. A few specimens from Peru (Amazonas) or less distinctly urceolate peri<strong>an</strong>th, sparsely to c<strong>an</strong>not be satisfactoraly inserted in one of these densely hairy; filaments longer th<strong>an</strong> the tepals, subspecies. They may represent <strong>an</strong>other subspefree. Pistillate inflorescences in fruit up to 20 cm cies. These collections (Kayap 201, Tunqui 217, long <strong><strong>an</strong>d</strong> 13 cm wide; peduncle 2-13 cm long, Huashikat 966, 1432 <strong><strong>an</strong>d</strong> 1482) largely match peduncle <strong><strong>an</strong>d</strong> br<strong>an</strong>ched sparsely to densely pub- the collections under subsp. tomentosa, but the erulous or also yellow-(sub)hirsute, often <strong>with</strong> arachnoid indument is almost lacking on most white, arachnoid hairs; flowers pedicellate (or parts, except for the main veins of the lamina subsessile), pedicel up to 0.5 cm long, in fruit up beneath <strong><strong>an</strong>d</strong> the petioles. Moreover, the hairs on to 1.5 cm long; peri<strong>an</strong>th 5-10 mm long, mostly the fruiting peri<strong>an</strong>th are much shorter, <strong><strong>an</strong>d</strong> the densely, yellowish, short-velutinous, sometimes apex of the leaves is short-acuminate. They are sparsely puberulous, or also <strong>with</strong> white, arach- provisionally placed under subsp. tomentosa. noid hairs; stigma (sub)peltate, ca. 1.5 mm diam. From the two syntype collections made by Fruiting peri<strong>an</strong>th black or purple, oblongoid to Martius, one is chosen as the lectotype collection. ellipsoid to ovoid, 1.5-2 x 0.8-1.5 cm, mostly densely, yellowish-puberulous to -velutinous or -subhirtellous. Key to the Subspecies of <strong>Pourouma</strong> tomentosa 1. Stipules hairy inside. .................................. ... ................. b. subsp. apiculata. 1. Stipules glabrous inside. 2. Leafy twigs yellow-hirsute (or glabrous); lamina 5-9-parted. ..................... c. subsp. persecta. 2. Leafy twigs puberulous or hirtellous; lamina entire or 3-5-parted. 3. Base of the lamina more or less deeply cordate; lamina 3-5-parted. ...... d. subsp. essequiboensis. 3. Base of the lamina rounded to truncate or subcordate; lamina usually entire. 4. Lamina 3-5-fid to -parted ............................................ a. subsp. tomentosa. 4. Lamina entire. 5. Apex of the lamina rounded to emarginate, (or short-acuminate in aberr<strong>an</strong>t specimens from Peru, Amazonas); heads of the staminate inflorescence 4-5 mm diam.; upper Amazon Basin. .......... ................................. a. subsp. tomentosa. 5. Apex of the lamina acuminate; heads of the staminate inflorescence 2-3 mm diam.; Gui<strong>an</strong>as, Amapa, <strong><strong>an</strong>d</strong> near M<strong>an</strong>aus. ............................. e. subsp. maroniensis. 13a. <strong>Pourouma</strong> tomentosa Miquel subsp. tomentosa. Fig. 79. <strong>Pourouma</strong> albistipulata Cuatrecasas, Bol. Soc. Venez. Ci. Nat. 17: 92. 1956. Type. Venezuela. Amazonas: Rio Guainia, Victorino, 22 Mar 1942 (9), LI. Williams 14833 (holotype, F; isotypes, F, G, NY, RB, S, US, VEN). Leafy twigs <strong><strong>an</strong>d</strong> petioles white-puberulous. Lamina entire, elliptic to ovate, sometimes (in Colombia <strong><strong>an</strong>d</strong> Ecuador) 3- or 5-parted, if entire, then often thickly coriaceous, + plicate, <strong><strong>an</strong>d</strong> apex rounded or emarginate (or sometimes short-acuminate), base rounded to truncate, or (in incised leaves) to subcordate; upper surface smooth; stipules outside <strong>with</strong> dense, white-arachnoid hairs, inside glabrous. Heads of the staminate inflorescences ca. 4-6 mm in diam. Pistillate flowers + diffusely distributed in the inflorescence, sometimes clustered in two groups; peduncle mostly up to 6(-7) cm long; peri<strong>an</strong>th <strong>with</strong>out dense, white, arachnoid indument, in fruit yellowish short-velutinous.
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Organization for Flora Neotropica C
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Copyright ? 1990 The New York Botan
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2 Flora Neotropica ABSTRACT Berg, C
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4 Flora Neotropica COUSSAPOA Engler
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6 Flora Neotropica Ni 3 4 5 FIG. 1.
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8 Flora Neotropica 1 2 3 4 5 6 7 a
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10 Flora Neotropica silicified oute
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12 Flora Neotropica v* r r'^S'9n 7
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1.trinervia *C.villosa Poeppig 6 En
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16 Flora Neotropica Table I Extreme
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18 Flora Neotropica 11. Lamina and
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20 Flora Neotropica anth; pistillat
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22 Flora Neotropica 66. Arachnoid h
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24 Flora Neotropica 2. Key to the C
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26 Flora Neotropica 3. Lateral vein
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28 Flora Neotropica 1. Interfloral
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30 10 Sep 1965 (st), Van Donselaar
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32 Flora Neotropica c. FIG. 12. Cou
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34 Flora Neotropica Key to the Subs
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36 Flora Neotropica 4b. Coussapoa a
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38 Flora Neotropica (MO); Dto. Anto
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40 Flora Neotropica FIG. 16. Coussa
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A' .. . FIG. 17. Coussapoa brevipes
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44 Flora Neotropica This species re
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46 Flora Neotropica 1 cm FIG. 20. C
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48 Flora Neotropica tion plane; pet
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50 Flora Neotropica This species ha
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52 Flora Neotropica I 1 cm FIG. 24.
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54 2 ~ ? I wo t :1cm Flora Neotropi
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56 Flora Neotropica G. 27. oussapoa
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58 Flora Neotropica to subsericeous
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60 cm1 2 I! 1 Flora Neotropica FIG.
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FIG. 31. Coussapoa latifolia: 1, le
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64 Flora Neotropica FG2 3,si1e tg t
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66 Flora Neotropica Distribution (F
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FIG.c3.m cousapoa anuenss 1 leves s
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70 Flora Neotropica berulous to hir
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FIG. 37. Coussapoa microcephala: ss
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74 Flora Neotropica ~.I. FIG. 38. C
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76 Flora Neotropica FIG. 39. Coussa
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78 Flora Neotropica ?~ ~~~~~~~~~~~~
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80 Specimens studied. MEXICO. TABAS
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82 Flora Neotropica ing the margin
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84 Flora Neotropica Coussapoa hirsu
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86 Specimens studied. BRAZIL. BAHIA
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FIG. 46. Coussapoa parvifolia: 1, l
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90 Flora Neotropica and Mocari, 150
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92 1cm Flora Neotropica FIG. 49. Co
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94 Flora Neotropica FIG. 51. Coussa
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2 1cm FIG. 52. Coussapoa trinervia:
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FG53 Cosaovanfla1leftwgwtstmntifoec
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100 Flora Neotropica FIG. 54. Couss
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102 Flora Neotropica 27336 (F, US).
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104 Flora Neotropica Loja, 1800 m,
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Coussapoa 107 FXG. 56. Coussapoafil
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Coussapoa FIG. 57. Coussapoa tolime
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Morphology ent revision, especially
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Morphology; Phenology; Pollination
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- Page 128 and 129: Pourouma125 restal "La Paragua," Ri
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- Page 132 and 133: Pourouma ~ I I MINISTERIO DE AGRICU
- Page 134 and 135: Pourouma 131 '~~~~~~~~~~~~~~~~~~~~~
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- Page 138 and 139: Pourouma 135 1953 (st), Steyermark
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- Page 142 and 143: Pourouma 139 This subspecies resemb
- Page 144 and 145: Pourouma 15520 (F, GH, NY), (9), Cu
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- Page 150 and 151: Pourouma 147 Rio Jurua, Marary, 8 O
- Page 152 and 153: Pourouma (U), 17 Feb 1980 (6), Berg
- Page 154 and 155: Pourouma ~~~~~~~~~~~~~~~~~~~~~~~~..
- Page 156 and 157: Pourouma 153 FG R.: 17. 411... . ur
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- Page 162 and 163: Pourouma159 Pourouma triloba Klotzs
- Page 164 and 165: Pourouma 161 between Rio Chicanan a
- Page 166 and 167: Pourouma 163 FIG. FLORA 76 Prom mn
- Page 170 and 171: + AJ ++ P O R O ULMA tomentosa. FIG
- Page 172 and 173: Pourouma 169 Varadouro do Morcego,
- Page 174 and 175: Pourouma qlq, _ j PIF P. t osantosa
- Page 176 and 177: Pourouma 173 465 (P); Trois Sauts,
- Page 178 and 179: Pourouma 175 I0711~~~~~~~~~~~~~~~~~
- Page 180 and 181: Pourouma 177 Specimens examined. VE
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- Page 184 and 185: Pourouma 181 the areoles; lateral v
- Page 186 and 187: Pourouma 183 Ak~~~~~~~~~~~~?IIk i.
- Page 188 and 189: Pourouma 185 I I STPE ,E Fn CL1 |SO
- Page 190 and 191: Pourouma 187 . . . FIG. 91. Pouroum
- Page 192 and 193: Pourouma 189 upper Rio Madre de Dio
- Page 194 and 195: Pourouma 191 bi f In i FIG. 92. Pou
- Page 196 and 197: Pourouma (BG), 9 Dec 1977 (d), Gent
- Page 198 and 199: Literature Cited 195 and distributi
- Page 200 and 201: Exsiccatae 197 e. subsp. chocoana (
- Page 202 and 203: Exsiccatae Dombey, J., s.n. (1-42).
- Page 204 and 205: Exsiccatae 201 Korning, J. et al.,
- Page 206 and 207: Exsiccatae 20); 15764 (1-40); 15791
- Page 208 and 209: Exsiccatae 205 Weberbauer, A., 3639
- Page 210 and 211: Index of Scientific Names 207 conto