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Cecropiaceae: Coussapoa and Pourouma, with an ... - CNCFlora

Cecropiaceae: Coussapoa and Pourouma, with an ... - CNCFlora

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<strong>Pourouma</strong> 155<br />

long, yellow-velutinous, <strong>with</strong> long, brown, pluri- mostly also on the smaller veins, occasionally<br />

cellular hairs; stigma peltate, ca. 1.5 mm diam., also on the leaf margin; lateral veins (6-)10-25<br />

yellow-puberulous. Fruiting peri<strong>an</strong>th ovoid to al- (-30) pairs, basal pair br<strong>an</strong>ched or sometimes (in<br />

most ellipsoid, 1.1-1.5 x 0.7-0.9 cm long, pu- obovate to elliptic leaves) unbr<strong>an</strong>ched, the basal<br />

berulous.<br />

part of these veins forming the basal part of the<br />

Distribution (Fig. 85). Peru (Loreto) <strong><strong>an</strong>d</strong> Brazil leaf margin (thus not separated from the margin<br />

(Amazonas, along Rio Javari); in non-inundated by mesophyll), reticulum <strong><strong>an</strong>d</strong> tertiary venation<br />

forest.<br />

(rather) pl<strong>an</strong>e beneath; petiole 3-22(-35) cm long,<br />

sparsely to densely white-puberulous (to -subve-<br />

Specimens examined. PERU. LORETO: Rio N<strong>an</strong>ay,<br />

Casario de Mish<strong>an</strong>a 3 Dec 1976 (9), Davidson 5203 lutinous) to yellow-hirsute, occasionally also <strong>with</strong><br />

(MO, U, US), 4 Dec 1976 (9), Rimachi 2725 (DUKE, long white, arachnoid hairs; stipules 3-16 cm long,<br />

F, NY); Iquitos, 3 Dec 1976 (9), Tessm<strong>an</strong>n 5364 (B, outside yellow(ish)-(sub)velutinous to -hirsute to<br />

S, US, type collection of P. phaeotricha). -(sub)sericeous, inside glabrous or sometimes<br />

BRAZIL. AMAZONAS: Rio Javari, Estirao do Ecuador,<br />

8 Aug 1973 (8), Lleras et al. P.17242 (F, INPA,<br />

sparsely hairy, caducous. Staminate inflores-<br />

MO, NY, U, US), 21 Oct 1976 (v), Pr<strong>an</strong>ce et al. 23966 cences up to 10 cm long <strong><strong>an</strong>d</strong> 5 cm wide; peduncle<br />

(G, INPA, NY, U).<br />

2-6 cm long, peduncle <strong><strong>an</strong>d</strong> br<strong>an</strong>ches whitish- to<br />

yellow-puberulous<br />

Vernacular names. Peru. Loreto: Sacha<br />

(to -subvelutionous) to -hiruvilla,<br />

tellous;flowers sessile or pedicellate, most of them<br />

uvilla.<br />

in<br />

The features of the staminate inflorescences<br />

globose heads, these 3-4 mm diam.; tepals<br />

0.5-1 mm<br />

<strong><strong>an</strong>d</strong> flowers <strong><strong>an</strong>d</strong> the pistillate flowers suggest close<br />

long, connate, forming a ? urceolate<br />

peri<strong>an</strong>th,<br />

relationships to P. heterophylla <strong><strong>an</strong>d</strong> P.<br />

hairy; filaments longer th<strong>an</strong> the periformica<strong>an</strong>th,<br />

free. Pistillate<br />

rum. However, P. phaeotricha lacks the swollen<br />

inflorescences in fruit up to<br />

18 cm<br />

petiole base.<br />

long <strong><strong>an</strong>d</strong> 10 cm wide; peduncle 3-12 cm<br />

The two collections from the<br />

long, peduncle <strong><strong>an</strong>d</strong> br<strong>an</strong>ches <strong>with</strong> indument sim-<br />

Brazili<strong>an</strong> b<strong>an</strong>k<br />

ilar to that of the staminate<br />

of Rio Javari have leaves <strong>with</strong> a scabrous<br />

inflorescence;flowers<br />

upper<br />

surface in<br />

10-30, in two clusters or more<br />

contrast to the other collections.<br />

diffusely distributed<br />

in the inflorescence; pedicel up to 1 cm, in<br />

fruit<br />

10. <strong>Pourouma</strong> mollis Trecul, Ann. Sci. Nat.<br />

up to 1.5 cm long, often already at <strong>an</strong>thesis<br />

Bot., <strong>with</strong> a<br />

Ser. 3, 8: 102, 1847; Miquel in Martius, Fl.<br />

strongly (+ capula-like) widened apex;<br />

bras. 4(1): 127. 1853; Berg, Fl. Suriname<br />

peri<strong>an</strong>th 0.5-1 cm long, yellow(ish)-(sub)ve-<br />

5(1):<br />

272, t. 15b. 1975. Type. French Gui<strong>an</strong>a.<br />

lutinous,<br />

Withapex<br />

? tuberculate; stigma peltate, ca.<br />

2 mm<br />

out locality, <strong>with</strong>out date (2), Leprieur s.n.<br />

diam., white-puberulous. Fruiting peri-<br />

(lec- <strong>an</strong>th black or<br />

totype, P. chosen by Berg, Fl. Suriname<br />

purplish, ovoid to ellipsoid or to<br />

5(1):<br />

272.<br />

oblongoid, 1-2 x 0.7-1.5 cm, usually<br />

1975;<br />

densely<br />

isolectotype, G).<br />

hairy.<br />

This species is very closely related to P. melinonii.<br />

We had considered recognizing the two<br />

taxa only at the subspecific level, but the lack of<br />

intermediates in areas where the two taxa grow<br />

together indicates that they are probably good<br />

species.<br />

Two allopatric subspecies c<strong>an</strong> be recognized.<br />

Tree, up to 30 m tall. Leafy twigs 4-10 mm<br />

thick, rather sparsely to densely white-puberu-<br />

lous (to -subvelutinous) to yellow-hirsute, <strong><strong>an</strong>d</strong><br />

<strong>with</strong> sparse to dense, purple to reddish-brown,<br />

pluricellular hairs, occasionally sparse, white,<br />

arachnoid hairs. Lamina entire, ovate to elliptic<br />

(or obovate), 6-25(-35) x 4-20(-30) cm or 3-5-<br />

lobed to -parted, ca. 10-25(-40) x 10-25(-40)<br />

cm, coriaceous (to subcoriaceous), apex acumi-<br />

nate, base acute, rounded to truncate, or sub-<br />

cordate; upper surface smooth, sometimes sca-<br />

brous, hairy on the main veins or subglabrous,<br />

occasionally yellow-hirsute, lower surface,<br />

sparsely to densely puberulous to hirtellous to<br />

subtomentose, often part of the hairs on the main<br />

veins appressed, arachnoid hairs in the areoles,<br />

Key to the Subspecies of<br />

<strong>Pourouma</strong> mollis<br />

1. Lamina usually entire; pistillate inflorescences<br />

<strong>with</strong> the flowers in two, more or less distinct<br />

clusters; Gui<strong>an</strong>as, Lower Amazon Basin, <strong><strong>an</strong>d</strong><br />

eastern Brazil. ................ a. subsp. mollis.<br />

1. Lamina usually 3-5 parted; pistillate inflorescences<br />

usually <strong>with</strong> the flowers diffusely distrib-

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