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English - Convention on Biological Diversity

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46<br />

COUNTRY STUDY FOR BIODIVERSITY OF THE REPUBLIC OF MACEDONIA<br />

as much as 500 t<strong>on</strong>nes. In the past few years it has been reduced to 70 t<strong>on</strong>nes, dropping<br />

to <strong>on</strong>ly 25 t<strong>on</strong>nes in 2002.<br />

The accelerated successi<strong>on</strong> of this lake ecosystem is evidenced by the appearance of<br />

the Calanoid copepod (Eudiaptomus gracilis), a typical representative of marsh<br />

ecosystems, which was recorded in Doyran Lake for the first time in 1995. In order to<br />

restore the disturbed envir<strong>on</strong>mental balance, efforts have been made to bring additi<strong>on</strong>al<br />

quantities of water to the lake, which is expected to improve the state of the biological<br />

communities within the lake ecosystem.<br />

The status of riverine ecosystems in the Republic of Maced<strong>on</strong>ia is also alarming.<br />

Almost all of the rivers are under great direct and/or indirect anthropogenic pressures.<br />

The situati<strong>on</strong> with the Vardar River, which is the major recipient of all types of<br />

wastewater (communal, industrial and agricultural), is the worst. The situati<strong>on</strong> with the<br />

other river ecosystems (Bregalnitsa, Crna, Lepenets, Pchinya, Zletovitsa etc.) is similar.<br />

Reservoirs have been built <strong>on</strong> some rivers, and these represent a sink for persistent<br />

substances (e.g., Kalimanci and Tikvesh Lakes). The reservoirs which provide drinking<br />

or industrial water (Mavrovitsa, Strezhevo, Turiya), although experiencing slight effects<br />

from natural eutrophicati<strong>on</strong>, have experienced a deteriorati<strong>on</strong> in quality in past years due<br />

to inappropriate fish stocking and exploitati<strong>on</strong>. Benthic communities in the riverine<br />

ecosystems are showing reduced abundance, which will ultimately lead to a decline in<br />

fish populati<strong>on</strong>s. Six out of the 20 endemic fish species within the Republic of<br />

Maced<strong>on</strong>ia (Ch<strong>on</strong>drostoma vardarense, Cobitis vardarensis, Gobio banarescui,<br />

Pachychil<strong>on</strong> maced<strong>on</strong>icum, Salmo pelag<strong>on</strong>icus and S. peristericus) are found in riverine<br />

ecosystems. Three of these are c<strong>on</strong>sidered to be globally threatened species.<br />

Wetland vegetati<strong>on</strong>, which used to develop over large areas of swamps and marshes<br />

within all the valleys of Maced<strong>on</strong>ia, experienced great changes under past drainage<br />

regimes which c<strong>on</strong>verted most of these ecosystems into arable land. In some of them,<br />

(e.g., M<strong>on</strong>ospitovo Marsh), numerous rare and endangered Algal taxa were formerly<br />

found.<br />

The relict wetland communities, which today appear mainly in a fragmentary state,<br />

are the most endangered. They develop <strong>on</strong> organic soils which are very suitable for<br />

growing early vegetable plants (Bansko) after drainage. Some which were present near<br />

natural lakes have been destroyed simply because they represent unwelcome marsh<br />

vegetati<strong>on</strong>. The most important wetland communities still extant are: assn. Caricetum<br />

elatae subassn. lysimachietosum (today <strong>on</strong>ly small fragments remain at Ohrid Lake near<br />

Studenchishte) assn. Cypero-Caricetum acutiformis (Gostivar), assn. Glycerietum<br />

maximae (Pelag<strong>on</strong>ia - village Chepigovo), assn. Mariscetum (Negortsi Spa), assn.<br />

Osmundo-Thelipteretum (Bansko), assn. Scirpo-Alopecuretum cretici (M<strong>on</strong>ospitovo<br />

Marsh) etc. Some of the wetlands which are still preserved are important in serving to<br />

explain the genesis of wetland vegetati<strong>on</strong> in the Republic of Maced<strong>on</strong>ia.<br />

Impacts to most of the swamps and marshes have caused a reducti<strong>on</strong> in the<br />

populati<strong>on</strong>s of all Amphibians, as well as individual species of other invertebrate and<br />

vertebrate groups. The most affected are: Water vole (Arvicola terrestris), Eurasian<br />

bittern (Botaurus stellaris), European p<strong>on</strong>d terrapin (Emys orbicularis), Otter (Lutra<br />

lutra), Balkan terrapin (Mauremys rivulata), Coypu (Myocastor coypus), Dice snake<br />

(Natrix tessellata), Miller’s water shrew (Neomys anomalus), Water shrew (N. fodiens),<br />

Muskrat (Ondatra zibethicus), Balkan spadefoot toad (Pelobates syriacus balcanicus),<br />

Eurasian Spo<strong>on</strong>bill (Platalea leucorodia), Greek marsh frog (Rana balcanica), Balkan<br />

stream frog (R. graeca), Marsh frog (R. ridibunda), Alpine newt (Triturus alpestris),

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