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Draft Black Sea Regional Transmission Project ESIA<br />

representing by Picea orientalis and Abies nordmanniana mixed with beech. Almost virgin<br />

dark coniferous forest occurs in Abastumani along the road to the observatory.<br />

Pine forests (Pinus kochiana) are usually developed on southern slopes of Meskheti, Adjara-<br />

Imereti and Trialeti ranges (Khintibidze, 1990). Pine forest has more limited distribution than<br />

spruce forests, although, pine frequently occurs in spruce forests on the northern slopes<br />

(Khintibidze, 1990). Pine forests on Erusheti and Tetrobi-Chobareti ranges (1800-2000<br />

meters above sea level) have little distinguished composition.<br />

Mountain steppes are peculiar to South Georgia, and cover the Javakheti volcanic plateau.<br />

Steppe vegetation is represented by different plant communities. Most characteristic species<br />

of polydominant grass-forb steppes are: Festuca ovina, F. sulcata, Stipa tirsa, S.<br />

pulcherrima, Bothriochloa ischaemum, Filipendula vulgaris, Falcaria vulgaris, Cruciata<br />

laevipes, Koeleria cristata, Medicago hemicycla, Phleum phleoides, Polygala anatolica,<br />

Thymus caucasicus, etc.<br />

There are also secondary meadows, principally on sites once occupied by primary forests.<br />

Like previous communities, these meadows are composed by the variants of polydominant<br />

grass-forb vegetation with participation of Agrostis planifolia, Alchemilla erythropoda,<br />

Brachypodium sylvaticum, Bromopsis variegata, Calamagrostis arundinacea, Centaurea<br />

salicifolia, Dactylis glomerata, Lotus caucasicus, Trifolium ambiguum, T. canescens, etc.<br />

From monodominant meadows can be mentioned communities with such dominant species<br />

as Nardus stricta (dzigviani in Georgian), Anemone fasciculata (frintiani), Agrostis planifolia<br />

(namikrefiani), Brachypodium sylvaticum (berseliani), Bromopsis variegata (shvrieliani), etc.<br />

(Kvachakidze, 1996).<br />

Subalpine Vegetation. The zubalpine zone is represented by krummholz, subalpine<br />

shrublands, tall herbaceous vegetation and polydominant subalpine meadows. Subalpine<br />

krummholz is represented by Betula litwinowii and B. pendula, Acer trautvetteri, Sorbus<br />

caucasigena, Salix caprea etc. Shrubland is composed of Caucasian Rhododendron -<br />

Rhododendron caucasicum, Vaccinium myrtillus, Empetrum caucasicum etc. Subalpine<br />

birch and maple forests are found on the northern slopes while pine forests are developed<br />

on the southern slopes at about 1800 to 1900 meters above sea level<br />

Only minor fragments of the former Javakheti subalpine forests survive, mostly on northern<br />

slopes of the high mountainous areas. These fragments are formed by species typical for the<br />

Caucasian subalpine forests, including Litvinov's birch (Betula litwinowii), mountain ash<br />

(Sorbus caucasigena), goat willow (Salix caprea), Bieberstein's rock currant (Ribes<br />

biebersteinii), alpine currant (Ribes alpinum), and in some areas European aspen (Populus<br />

tremula) and others. Litvinov's birch and mountain ash form communities that cover areas of<br />

rocky relief.<br />

Tall herbaceous vegetation is composed of 3-4 meter-high herbs, mainly dicots<br />

(Nakhutsrishvili, 1999). Typical species forming subalpine tall herbaceous vegetation are as<br />

follows: Anemone fasciculata, Geranium ibericum, G. platypetalum, G. psilostemon, G.<br />

ruprechtii, Scabiosa caucasica, Senecio rhombifolius, Stachys macrantha, Campunala<br />

latifolia, Cephalaria gigantea, Doronicum macrophyllum, Aconitum nasutum, Gadellia<br />

lactiflora, Delphinium flexuosum, Heracleum wilhelmsii, Grossheimia macrocephala, Lilium<br />

szovitsianum, etc.<br />

Subalpine grass and grass forb meadows are found in the subalpine forest complexes.<br />

Grass meadows are formed by Festuca ovina, F. woronowii, Bromopsis variegata,<br />

Calamagrostis arundinacea. These species form coenoses both independently and in codominance.<br />

The subalpine meadows occur above the subalpine forest zone, at the altitudes<br />

of 2100 to 2200 meters above sea level<br />

74

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