472ACANTHACEAEin corolla tube; thecae ca. 1 mm, pilose at tip. Ovary oblong;style sparingly hirsute. Capsule ca. 4 mm. Seeds subcircular inoutline.Taiwan [Philippines].No specimens of Lepidagathis secunda have been seen, and publisheddescriptions are inadequate to evaluate the status of either thisspecies or its occurrence in Taiwan. Information above is based entirelyon that provided by C. F. Hsieh and T. C. Huang (Fl. Taiwan, ed. 2, 4:673. 1998).7. Lepidagathis stenophylla C. B. Clarke ex Hayata, J. Coll.Sci. Imp. Univ. Tokyo 30(1): 214. 1911.柳 叶 鳞 花 草 liu ye lin hua caoHerbs to 40 cm tall, suffrutescent, often ± anisophyllous.Stems 4-angled to 4-angled and sulcate, subglabrous or evenlyto ± bifariously pubescent. Petiole (1–)5–12 mm; leaf bladelinear to linear-lanceolate, 3.5–11 × 0.3–1.9 cm, (4.6–)6.5–16 ×as long as wide, both surfaces subglabrous or pubescent,secondary veins 4 or 5 on each side of midvein, base narrowedand decurrent onto petiole, margin shallowly sinuate, apexacute to acuminate. Spikes elongate, 1.3–3.8 cm, secund; bractslanceolate to lanceolate-ovate, ca. 5 × 1–1.5 mm, abaxially andmarginally pubescent with non-glandular trichomes, 1-veined,apex aristate; bracteoles similar to bracts except usually smallerand narrower. Calyx 5.5–8.2 mm, pubescent with non-glandulartrichomes; posterior lobe ovate to oblong-lanceolate, 5–7.5 ×1.1–1.3 mm, 3-veined; lateral lobes lanceolate, 4.5–5.7 × 0.7–0.8 mm; anterior lobes connate at base for 20%–55% of theirlength, distinct segments narrowly lanceolate and 3–6 mm.Corolla white, ca. 5 mm, outside pubescent; lower lip 2–2.3mm, lobes 1.5–1.6 × 0.6–0.8 mm. Capsule 4.5–5.5 mm, pubescent± throughout or only distally. Seeds subcircular in outline,1–1.8 mm in diam. Fl. and fr. Dec.● Forest trails; 200–400 m. Hong Kong, Taiwan.31. CHROESTHES Benoist, Bull. Mus. Natl. Hist. Nat. 33: 107. 1927.色 萼 花 属 se e hua shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielShrubs, subisophyllous or anisophyllous. Leaves opposite, petiolate; leaf blade margin entire. Inflorescences terminal thyrses;bracts and bracteoles greenish. Calyx unequally 5-lobed; posterior lobe largest, 2 lateral lobes smaller than 2 anterior lobes. Corollatube basally cylindric and expanded distally into a throat; limb 2-lipped, lower lip 3-lobed, upper lip 2-lobed; lobes quincuncial inbud. Stamens 4, not connate, posterior pair shorter than anterior pair, inserted at base of throat; anthers 2-thecous; thecae parallel,inserted at different heights, dorsally pubescent, base spurred. Ovary with 2 ovules per locule; style basally sparsely pubescent.Capsule stipe absent or barely present; retinacula present. Seeds compressed, brownish pubescent.Three species: China, Laos, Malaysia, Myanmar, Thailand, Vietnam; one species in China.1. Chroesthes lanceolata (T. Anderson) B. Hansen, Nordic J.Bot. 3: 209. 1983.色 萼 花 se e huaAsystasia lanceolata T. Anderson, J. Linn. Soc., Bot. 9:524. 1867; A. kerrii Craib; A. silvicola W. W. Smith; Chroesthespubiflora Benoist; C. racemiflora Bremekamp; C. racemifloravar. glabrior Bremekamp; C. silvicola (W. W. Smith) E.Hossain.Shrubs 0.5–3 m tall, anisophyllous. Stems terete, slender,rarely branched, glabrous. Petiole 1–2.5 cm; leaf blade ellipticto oblanceolate to lanceolate, 10–16 × 3–7 cm, both surfacesglabrous, secondary veins 6–9 on each side of midvein, basecuneate, margin entire or subsinuate, apex acuminate. Thyrses3–7 cm; cymes sessile, 1–3-flowered; bracts elliptic to broadlylanceolate, 3–9 × 1–3 mm, gland-tipped pubescent, apex acute;bracteoles narrowly elliptic to broadly lanceolate, 4–9 × 0.7–1.2mm. Pedicel 1–5 mm. Calyx 1–1.6 cm, outside gland-tippedpubescent; posterior lobe lanceolate, ovate, or subelliptic; laterallobes linear-lanceolate; anterior lobes connate to 2/3 theirlength. Corolla white with pink or purple spots, ca. 2.5 cm, outsidepubescent; tube basal portion ca. 9 mm, throat ca. 1.5 cm;lower lip 3-lobed; upper lip 2-lobed. Stamens included in throat;filaments 1–1.2 cm, glabrous; anther thecae 2.1–2.3 mm, pubescentat apex and along sides, basal spur pointed. Ovary apexpubescent; style ca. 2.5 cm. Capsule subellipsoid to obovoid,1.2–1.6 cm, glabrous or only at apex pubescent, 4-seeded.Seeds subcircular in outline. Fl. Feb–Mar, fr. May–Jul.Forests; (200–)800–1400 m. Guangxi, S Yunnan [Laos, Myanmar,Thailand, Vietnam].32. GYMNOSTACHYUM Nees in Wallich, Pl. Asiat. Rar. 3: 76, 106. 1832.裸 柱 草 属 luo zhu cao shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielCryptophragmium Nees; Sarcanthera Rafinesque.Herbs, perennial [or shrubs], with cystoliths. Leaves cauline or subradical, opposite; leaf blade margin usually entire. Inflorescencesaxillary or terminal, spikes, racemes, or panicles, often secund; bracts and bracteoles shorter than calyx. Calyx 5-lobed nearlyto base; lobes subequal. Corolla tube longer than limb, basal portion narrowly cylindric, ± expanded distally into a throat; limb 2-lipped, lower lip 3-lobed, upper lip emarginate to 2-lobed; lobes ascending cochlear in bud. Stamens 2, included in or slightly
ACANTHACEAE 473exserted from corolla tube; anther thecae 2, equal or subequal in size, parallel, one or both mucronate at base; staminodes absent.Ovary with 3 to many ovules per locule; stigma 2-lobed, lobes compressed. Capsule without a stipe, linear, 4-angled, many seeded;retinacula present. Seeds compressed, covered with hygroscopic trichomes.About 30 species: tropical Asia; three species (two endemic) in China.1a. Stems very short; leaves in a basal rosette ............................................................................................................. 1. G. subrosulatum1b. Stems elongate; leaves cauline.2a. Inflorescences axillary .................................................................................................................................................. 2. G. listeri2b. Inflorescences terminal .............................................................................................................................................. 3. G. sinense1. Gymnostachyum subrosulatum H. S. Lo, Acta Phytotax.Sin. 17(4): 86. 1979.矮 裸 柱 草 ai luo zhu caoGymnostachyum kwangsiense H. S. Lo.Herbs 20–40 cm tall. Stems short, hirsute. Leaves in abasal rosette; petiole 2–9(–14) cm; leaf blade suborbicular tobroadly ovate, 5–16 × 3.5–11 cm, papery, abaxially subglabrousbut midvein and secondary veins hirsute, adaxially slightlyscabrous, secondary veins 7–9 on each side of midvein, baseslightly cordate to rounded, margin entire or slightly sinuate,apex rounded to acute. Thyrses 10–35 cm, with 1–4 flowers percyme; peduncle 8–19 cm; bracts subulate, ca. 2.5 mm, pilose,apex acute; bracteoles similar to bracts. Calyx 2.5–3 mm; lobessubulate, pilose. Corolla ca. 1.5 cm; lower lip patent, 3-lobed,lobes oblong-ovate and ca. 3 × 2 mm; upper lip triangular, ca. 5mm, erect, 2-dentate. Stamens exserted; filaments 7–8 mm; antherthecae linear, parallel. Ovary 2-locular, with 4 ovules perlocule; style 1.3–1.9 cm. Capsule linear, 1.3–1.6 cm, 8-seeded.Seeds ovate in outline, ca. 1 × 0.8 mm. Fl. Apr–May, fr. Jun–Sep.● Limestone hills; 200–600 m. Guangxi.2. Gymnostachyum listeri Prain, J. Asiat. Soc. Bengal, Pt. 2,Nat. Hist. 69: 171. 1900.云 南 裸 柱 草 yun nan luo zhu caoCryptophragmium tonkinense Benoist; Parajusticia petelotiiBenoist.Herbs to 50 cm tall, caulescent, isophyllous. Stemsbranched, 4-angled, pubescent. Leaves cauline; petiole 5–12mm, pubescent; leaf blade elliptic to obovate-elliptic, (4.5–)6–13(–17) × 2.1–5.5 cm, abaxially white pubescent, adaxially glabrousexcept for trichomes along midvein, secondary veins 11–14 on each side of midvein and abaxially prominently curved,tertiary veins inconspicuous, base asymmetric or symmetric and± decurrent onto petiole, margin entire or obscurely sinuate,apex acute to acuminate. Spikes 2–6 cm, secund; peduncle 0.5–1.5 cm; rachis pubescent and gland-tipped pilose; bracts linear,ca. 3 × 0.3 mm, pubescent; bracteoles linear-triangular, ca. 1.5 ×0.5 mm, pubescent. Calyx 0.1–1.1 cm; lobes linear, 8–10 × ca.0.5 mm, outside pubescent, inside glabrous, margin glandtippedpilose. Corolla purple, 1.6–2 cm, pubescent; tube basallycylindric and ca. 2.5 cm wide for ca. 8 mm, throat abruptlybending and widening to 5 mm; lower lip 4–5 mm, 3-lobed,lobes oblong and ca. 3 × 1.5 mm; upper lip 3–4 mm, 2-lobed,lobes broadly ovate and ca. 2 × 2 mm. Staminal filaments ca.4 mm, glabrous; anther thecae oblong, ca. 5 × 1 mm, pilose,spurred at base. Ovary ca. 3 × 1 mm, glabrous; style ca. 1.5 cm,villous. Capsule not seen. Fl. Apr–May.Limestone hills; 200–600 m. Guangxi, Yunnan [Bangladesh, Vietnam].3. Gymnostachyum sinense (H. S. Lo) H. Chu, Bull. Bot. Res.,Harbin 11(1): 48. 1991.华 裸 柱 草 hua luo zhu caoAndrographis sinensis H. S. Lo, Bull. Bot. Res., Harbin1(4): 103. 1981.Herbs to 1 m tall. Stems purplish black, terete, glabrous;young shoots 4-angled, pubescent at nodes. Petiole 2–4 mm;leaf blade ovate to broadly ovate, 1.5–4 × 1–1.8 cm, thicklypaper, both surfaces glabrous, secondary veins ca. 6 on eachside of midvein, base cuneate, margin entire or subentire, apexacute to sometimes caudate. Spikes terminal, few flowered; peduncleca. 1 cm, subglabrous; bracts linear to linear-lanceolate,1.5–2 mm; bracteoles similar to bracts. Calyx ca. 7 mm, glabrous;lobes lanceolate-linear, apex acuminate. Corolla white,ca. 1 cm; lower lip ca. 4 mm, 3-lobed, lobes ovate and ca. 0.5 ×0.5 mm; upper lip subtriangular, ca. 3.5 mm, 2-lobed. Staminalfilaments ca. 2.5 mm, glabrous; anther thecae linear, ca. 3 mm,spurred at base. Ovary ellipsoid, ca. 3 mm, glabrous, ovules 5or 6 per locule; style ca. 7 mm. Capsule narrowly cylindric, 1–1.5 cm, glabrous. Seeds subglobose, ca. 1 × 1 mm, pubescent.Fl. Aug.● Limestone hills. Guangxi.33. ANDROGRAPHIS Wallich ex Nees in Wallich, Pl. Asiat. Rar. 3: 77, 116. 1832.穿 心 莲 属 chuan xin lian shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielHaplanthoides H. W. Li; Haplanthus Nees.Herbs or subshrubs, with cystoliths. Leaf blade margin entire. Inflorescences terminal or axillary, panicles, racemes, or sometimesspikes or dense clusters; bracts present; bracteoles small [or absent]. Calyx deeply 5-lobed; lobes narrow, equal or subequal.Corolla tubular to funnelform at base; limb 2-liped; lower lip 3-lobed; upper lip entire or 2-lobed; lobes ascending cochlear in bud.
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ACANTHACEAE爵 床 科 jue chuang k
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ACANTHACEAE 37122b. Anthers 2-theco
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ACANTHACEAE 37311a. Inflorescences
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ACANTHACEAE 375lary or terminal, 2-
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ACANTHACEAE 377Two species: China,
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ACANTHACEAE 379Vines, herbaceous. S
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ACANTHACEAE 3817. STROBILANTHES Blu
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ACANTHACEAE 383Type 10. S. tetraspe
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ACANTHACEAE 38525b. Plants strongly
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ACANTHACEAE 38775a. Calyx lobes lin
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ACANTHACEAE 389115b. Corolla outsid
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ACANTHACEAE 391times rosette-formin
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ACANTHACEAE 393576. 1784; Acanthopa
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ACANTHACEAE 395with a whitish tube
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ACANTHACEAE 397Herbs 30-80 cm tall,
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ACANTHACEAE 399Strobilanthes petiol
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ACANTHACEAE 401silky tomentose and
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ACANTHACEAE 4036 × 1.2-1.5 mm. Cor
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ACANTHACEAE 405acute to shortly acu
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ACANTHACEAE 407Subshrubs to 20 cm t
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ACANTHACEAE 40910 × ca. 3 mm, apic
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ACANTHACEAE 4111b. Stems, petioles,
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ACANTHACEAE 413woody. Leaves subses
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ACANTHACEAE 415gins white ciliolate
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ACANTHACEAE 417long acuminate; brac
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ACANTHACEAE 419retaining style; tub
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