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472ACANTHACEAEin corolla tube; thecae ca. 1 mm, pilose at tip. Ovary oblong;style sparingly hirsute. Capsule ca. 4 mm. Seeds subcircular inoutline.Taiwan [Philippines].No specimens of Lepidagathis secunda have been seen, and publisheddescriptions are inadequate to evaluate the status of either thisspecies or its occurrence in Taiwan. Information above is based entirelyon that provided by C. F. Hsieh and T. C. Huang (Fl. Taiwan, ed. 2, 4:673. 1998).7. Lepidagathis stenophylla C. B. Clarke ex Hayata, J. Coll.Sci. Imp. Univ. Tokyo 30(1): 214. 1911.柳 叶 鳞 花 草 liu ye lin hua caoHerbs to 40 cm tall, suffrutescent, often ± anisophyllous.Stems 4-angled to 4-angled and sulcate, subglabrous or evenlyto ± bifariously pubescent. Petiole (1–)5–12 mm; leaf bladelinear to linear-lanceolate, 3.5–11 × 0.3–1.9 cm, (4.6–)6.5–16 ×as long as wide, both surfaces subglabrous or pubescent,secondary veins 4 or 5 on each side of midvein, base narrowedand decurrent onto petiole, margin shallowly sinuate, apexacute to acuminate. Spikes elongate, 1.3–3.8 cm, secund; bractslanceolate to lanceolate-ovate, ca. 5 × 1–1.5 mm, abaxially andmarginally pubescent with non-glandular trichomes, 1-veined,apex aristate; bracteoles similar to bracts except usually smallerand narrower. Calyx 5.5–8.2 mm, pubescent with non-glandulartrichomes; posterior lobe ovate to oblong-lanceolate, 5–7.5 ×1.1–1.3 mm, 3-veined; lateral lobes lanceolate, 4.5–5.7 × 0.7–0.8 mm; anterior lobes connate at base for 20%–55% of theirlength, distinct segments narrowly lanceolate and 3–6 mm.Corolla white, ca. 5 mm, outside pubescent; lower lip 2–2.3mm, lobes 1.5–1.6 × 0.6–0.8 mm. Capsule 4.5–5.5 mm, pubescent± throughout or only distally. Seeds subcircular in outline,1–1.8 mm in diam. Fl. and fr. Dec.● Forest trails; 200–400 m. Hong Kong, Taiwan.31. CHROESTHES Benoist, Bull. Mus. Natl. Hist. Nat. 33: 107. 1927.色 萼 花 属 se e hua shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielShrubs, subisophyllous or anisophyllous. Leaves opposite, petiolate; leaf blade margin entire. Inflorescences terminal thyrses;bracts and bracteoles greenish. Calyx unequally 5-lobed; posterior lobe largest, 2 lateral lobes smaller than 2 anterior lobes. Corollatube basally cylindric and expanded distally into a throat; limb 2-lipped, lower lip 3-lobed, upper lip 2-lobed; lobes quincuncial inbud. Stamens 4, not connate, posterior pair shorter than anterior pair, inserted at base of throat; anthers 2-thecous; thecae parallel,inserted at different heights, dorsally pubescent, base spurred. Ovary with 2 ovules per locule; style basally sparsely pubescent.Capsule stipe absent or barely present; retinacula present. Seeds compressed, brownish pubescent.Three species: China, Laos, Malaysia, Myanmar, Thailand, Vietnam; one species in China.1. Chroesthes lanceolata (T. Anderson) B. Hansen, Nordic J.Bot. 3: 209. 1983.色 萼 花 se e huaAsystasia lanceolata T. Anderson, J. Linn. Soc., Bot. 9:524. 1867; A. kerrii Craib; A. silvicola W. W. Smith; Chroesthespubiflora Benoist; C. racemiflora Bremekamp; C. racemifloravar. glabrior Bremekamp; C. silvicola (W. W. Smith) E.Hossain.Shrubs 0.5–3 m tall, anisophyllous. Stems terete, slender,rarely branched, glabrous. Petiole 1–2.5 cm; leaf blade ellipticto oblanceolate to lanceolate, 10–16 × 3–7 cm, both surfacesglabrous, secondary veins 6–9 on each side of midvein, basecuneate, margin entire or subsinuate, apex acuminate. Thyrses3–7 cm; cymes sessile, 1–3-flowered; bracts elliptic to broadlylanceolate, 3–9 × 1–3 mm, gland-tipped pubescent, apex acute;bracteoles narrowly elliptic to broadly lanceolate, 4–9 × 0.7–1.2mm. Pedicel 1–5 mm. Calyx 1–1.6 cm, outside gland-tippedpubescent; posterior lobe lanceolate, ovate, or subelliptic; laterallobes linear-lanceolate; anterior lobes connate to 2/3 theirlength. Corolla white with pink or purple spots, ca. 2.5 cm, outsidepubescent; tube basal portion ca. 9 mm, throat ca. 1.5 cm;lower lip 3-lobed; upper lip 2-lobed. Stamens included in throat;filaments 1–1.2 cm, glabrous; anther thecae 2.1–2.3 mm, pubescentat apex and along sides, basal spur pointed. Ovary apexpubescent; style ca. 2.5 cm. Capsule subellipsoid to obovoid,1.2–1.6 cm, glabrous or only at apex pubescent, 4-seeded.Seeds subcircular in outline. Fl. Feb–Mar, fr. May–Jul.Forests; (200–)800–1400 m. Guangxi, S Yunnan [Laos, Myanmar,Thailand, Vietnam].32. GYMNOSTACHYUM Nees in Wallich, Pl. Asiat. Rar. 3: 76, 106. 1832.裸 柱 草 属 luo zhu cao shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielCryptophragmium Nees; Sarcanthera Rafinesque.Herbs, perennial [or shrubs], with cystoliths. Leaves cauline or subradical, opposite; leaf blade margin usually entire. Inflorescencesaxillary or terminal, spikes, racemes, or panicles, often secund; bracts and bracteoles shorter than calyx. Calyx 5-lobed nearlyto base; lobes subequal. Corolla tube longer than limb, basal portion narrowly cylindric, ± expanded distally into a throat; limb 2-lipped, lower lip 3-lobed, upper lip emarginate to 2-lobed; lobes ascending cochlear in bud. Stamens 2, included in or slightly

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