440ACANTHACEAEEranthemum shweliense W. W. Smith, Notes Roy. Bot.Gard. Edinburgh 10: 176. 1918.Shrubs 1–2 m tall. Stems woody, grayish, hispid whenyoung. Petiole 3–5 mm, densely brownish hirsute; leaf bladelanceolate, ovate-lanceolate, or elliptic, 2.5–5.5 × 1–2 cm, abaxiallysparsely pilose or glabrous but densely brown hairy alongveins, adaxially sparsely pilose along veins, base cuneate anddecurrent onto petiole, margin entire, apex obtuse to acute. Inflorescencesterminal, 7–15 cm; rachis densely yellowish pilose,with flowers fascicled at each node; bracts linear to linearlanceolate,ca. 3 mm, pilose; bracteoles subulate, ca. 2 mm.Pedicel ca. 2 mm, pilose. Calyx ca. 5 mm; lobes linear-oblong,pilose, apex acute. Corolla white with violet spot on 2 lobes;tube ca. 2.2 cm, outside and inside pilose; lobes 5, ovate,1.8–1.9 cm. Stamens exserted. Ovary glabrous; style basallysparsely pilose; stigma slightly 2-cleft. Capsule clavate, ca. 2.5cm. Seeds ovate in outline, ca. 3 mm, verrucose, glabrous. Fl.Feb–May, fr. Jun–Aug.● Thickets, forest margins; 1200–1800 m. Yunnan.4. Pseuderanthemum teysmannii (Miquel) Ridley, Fl. Malay.Penin. 2: 592. 1923 [“teysmanni”].红 河 山 壳 骨 hong he shan ke guStrobilanthes teysmannii Miquel, Fl. Ned. Ind. 2: 799.1858 [“teysmanni”].Shrubs. Stems reddish brown, 4-angled, glabrous. Petioleto 2.5 cm; leaf blade ovate, 3.5–9.5 × 1.5–4.5 cm, abaxially pubescentalong veins, adaxially sparsely pubescent along midvein,secondary veins 8–10 on each side of midvein, basebroadly cuneate and decurrent onto petiole, margin subentire,apex acuminate. Racemes terminal or axillary; rachis with flowers0.5–1.5 cm apart, yellowish brown pubescent; bracts triangular,2–3 mm, margin ciliate; bracteoles similar to bracts. Pedicel0–4 mm, pilose. Calyx ca. 5 mm, pubescent; lobes linear,slightly unequal. Corolla pale purple, ca. 3.5 cm; tube ca. 2.5cm; limb 5-lobed, slightly 2-lipped. Stamens inserted at corollathroat; anthers as long as filaments. Ovary pilose; style ca. 2cm, slightly white villous; stigma 2-cleft. Capsule clavate, ca.2.3 cm, sparsely pubescent. Seeds ovate in outline, ca. 3 × 3mm, verrucose, glabrous. Fl. Apr–Jun, fr. Aug.Thickets; 100–300 m. Yunnan [Indonesia, S Thailand].5. Pseuderanthemum coudercii Benoist, Notul. Syst. (Paris)5: 111. 1936 [“couderci”].狭 叶 钩 粉 草 xia ye gou fen caoSubshrubs or perennial herbs, 30–50 cm tall. Stems subtereteto 4-angled, pubescent, glabrescent. Petiole ca. 5 mm; leafblade oblong, lanceolate, or linear, 3–11 × 0.2–2.5 mm, bothsurfaces glabrous, secondary veins 6–12 on each side of midvein,base cuneate, margin entire or slightly undulate, apex acuminateto rarely acute. Thyrses terminal, to 20 cm; rachis pubescent;dichasia 1–4-flowered; basal bracts 1–3, large, leaflike,longer than calyx; apical bracts linear, shorter than calyx, villous;bracteoles linear, villous. Calyx 5–6 mm; lobes linear, villous.Corolla light purplish red, slightly pubescent; tube ca. 3cm; lower lip ca. 1.2 cm, lobes ovate to ovate-elliptic; upperlip 2-lobed, lobes oblong. Stamens 2; filaments connate atbase; staminodes 2, smaller than stamens. Capsule ca. 1.8 cm,sparsely pubescent. Seeds obovate in outline, ca. 3.5 mm, tuberculate-punctate,glabrous. Fl. Jun–Aug, fr. Sep.Forests, streamsides; 100–400 m. Hainan [Cambodia].6. Pseuderanthemum haikangense C. Y. Wu & H. S. Lo, Fl.Hainan. 3: 595. 1974.海 康 钩 粉 草 hai kang gou fen caoShrubs 40–100 cm tall. Stems terete, glabrous; bark strawyellow.Petiole 0.5–1.5 cm; leaf blade elliptic-orbicular, oblong,ovate, or rarely lanceolate-elliptic, 5–11.5 × 2–3.5 cm,both surfaces glabrous, secondary veins 5–7 on each side ofmidvein, base broadly cuneate to rounded, margin entire or inconspicuouslyundulate, apex shortly acuminate to caudate. Racemesterminal or rarely in apical leaf axils, to 30 cm; bractsnarrowly triangular, 2–3 mm; bracteoles 1–2 mm. Pedicel 0–3mm. Calyx ca. 5 mm; lobes linear-lanceolate, apex acuminate.Corolla white or pink and with red spots on lobes of lower lips,ca. 4 cm, pilose; tube basally ca. 1.5 mm wide for ca. 3 cm;lower lip 3-lobed, lobes narrowly elliptic, ca. 10 × 4 mm, andapex obtuse; upper lip 2-lobed, lobes elliptic, ca. 10 × 6 mm,and apex obtuse. Stamens 2; filaments ca. 2 mm; anther thecaeoblong, ca. 1.5 mm; staminodes ca. 1.3 mm. Ovary pilose; styleca. 3 cm, slightly pubescent; stigma slightly 2-cleft. Capsuleclavate, 1.9–2 cm, pubescent. Seeds broadly ovate in outline,ca. 3.5 mm, tuberculate-punctate. Fl. Jun–Aug, fr. Sep.● Forests; 200–900 m. Guangdong, Hainan, Yunnan.7. Pseuderanthemum latifolium (Vahl) B. Hansen, Nordic J.Bot. 9: 213. 1989.山 壳 骨 shan ke guJusticia latifolia Vahl, Symb. Bot. 2: 4. 1791; Antheliacanthusmicranthus Ridley; Eranthemum palatiferum (Wallich)Nees; J. palatifera Wallich; Pseuderanthemum palatiferum(Wallich) Radlkofer.Herbs to 1 m tall, perennial. Stems 4-angled, pubescent.Petiole 1–2.5 mm; leaf blade elliptic, 11.5–12 × 3.5–6 cm,abaxially glabrous except villous on midvein, adaxially glabrous,secondary veins 5 or 6 on each side of midvein, basecuneate, margin subentire to slightly undulate-crenulate, apexacuminate. Racemes to 30 cm; rachis pubescent, usually with 3-flowered clusters at nodes; bracts triangular, 4–6 × ca. 2 mm,pubescent; bracteoles similar to bracts. Pedicel 0.5–2 mm. Calyx5–6 mm, outside pubescent; lobes linear, 4.5–5.5 × ca. 1mm. Corolla light purple with purple dots on lower lip, 2–3 cm;tube linear, 1.5–2.5 cm; lobes 3–4 × ca. 3 mm. Style basally pubescent.Capsule ca. 2.5 cm, pubescent. Seeds ovate in outline,ca. 4 × 4 mm, rugose. Fl. Apr–Jun, fr. Aug–Sep.Forests; 100–1600 m. Guangdong, Guangxi, Hainan, Yunnan[Cambodia, India, Laos, Malaysia, Myanmar, Thailand, Vietnam].
ACANTHACEAE 44116. MACKAYA Harvey, Thes. Cap. 1: 8. 1859, nom. cons., not Arnott (1838),nor Mackaia Gray (1821).太 平 爵 床 属 tai ping jue chuang shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielHerbs, perennial, with cystoliths. Leaves petiolate. Inflorescences axillary or terminal, secund racemes; bracts linear-lanceolate[or narrowly deltoid]; bracteoles 2, borne at base of pedicel. Calyx much shorter than corolla tube, 5-lobed almost to base; lobeslinear-lanceolate. Corolla subactinomorphic; tube basally cylindric, campanulate toward apex; limb 2-lipped, lower lip 3-lobed,upper lip 2-lobed; lobes suborbicular [ovate or oblong], ascending cochlear in bud. Stamens 2, inserted at base of campanulate portionof corolla tube, subincluded; anthers 2-thecous; thecae oblong, dorsally pubescent; staminodes 2, lacking anthers. Disk obsolete.Ovary subellipsoid, with ovules 2 per locule; style subglabrous or pubescent near base; stigma 2-lobed. Capsule (not seen forMackaya tapingensis) stipitate, 2- or 4-seeded; retinacula present. Seeds (not seen for M. tapingensis) somewhat rugose.Three species: disjunctly distributed in Himalayan region and South Africa; one species in China.1. Mackaya tapingensis (W. W. Smith) Y. F. Deng & C. Y. Wu,Novon 19: 308. 2009.太 平 爵 床 tai ping jue chuangEranthemum tapingense W. W. Smith, Notes Roy. Bot.Gard. Edinburgh 10: 177. 1918; Pseuderanthemum tapingense(W. W. Smith) C. Y. Hu & H. S. Lo.Herbs to 1 m tall, perennial. Stems subterete, glabrous orsubglabrous. Petiole 2–3.5 cm, sparsely puberulent or glabrescent;leaf blade lanceolate to oblong-lanceolate, 12–17 × 3–5(–7) cm, membranous, both surfaces glabrous or glabrescent,abaxially paler, adaxially dark green, secondary veins 8–12 oneach side of midvein, base cuneate, margin entire to undulate,apex acuminate. Inflorescences axillary or terminal, racemes,7–20 cm; peduncle sparsely pilose or glabrescent; bracts linearlanceolate,2–3 mm, pilose or glabrescent; bracteoles linear, ca.1 mm. Pedicel 2–3 mm. Calyx 6–8 mm; lobes linear-lanceolate,3–4 × 1.5–2 mm, glabrous, apex acuminate. Corolla rose, 3–4cm, outside glabrous; tube basally cylindric for ca. 1 cm; lobessuborbicular, 7–12 mm, subequal, margin dentate. Staminal filaments6–12 mm, pilose; anther thecae 4–5 mm, muticous atbase; staminodes 1–4 mm. Ovary glabrous; style ca. 1.8 cm, pubescent.Capsule not seen. Fl. Oct–Jan.Forests; 600–1800 m. Yunnan [Myanmar].17. ISOGLOSSA Oersted, Vidensk. Meddel. Dansk Naturhist. Foren. Kjøbenhavn1854: 155. 1854, nom. cons.叉 序 草 属 cha xu cao shuHu Jiaqi ( 胡 嘉 琪 Hu Chia-chi), Deng Yunfei ( 邓 云 飞 ); Thomas F. DanielChingiacanthus Handel-Mazzetti; Ecteinanthus T. Anderson; Leda C. B. Clarke (1908), not Bory (1822) [Algae]; RhytiglossaNees; Strophacanthus Lindau.Shrubs or herbs, with cystoliths, isophyllous or slightly anisophyllous. Leaves petiolate. Inflorescences terminal and/or sometimesaxillary, open panicles or thyrses; bracts small, shorter than calyx. Calyx deeply 5-lobed, lobes similar. Corolla funnel-shaped,2-lipped; lower lip 3-lobed; upper lip shortly 2-lobed, lacking a stylar furrow (i.e., erugulate); lobes ascending cochlear in bud.Stamens 2; anthers 2-thecous; thecae equal or unequal, parallel (to perpendicular), subequally to unequally inserted, base muticous;staminodes absent; pollen 2-pororate [elsewhere also 3-colporate and 6-pseudocolpate]. Ovary with 2 ovules per locule. Capsuleclavate with a solid sterile basal stalk, 4-seeded; retinacula present; septa with attached retinacula remaining attached to inner wall ofmature capsule. Seeds subdiscoid.About 50 species: tropical Africa and Asia; two species (one endemic) in China.1a. Calyx lobes subulate, widest at base; inflorescence long branched, widely paniculate ................................................... 1. I. collina1b. Calyx lobes spatulate-oblanceolate, widest above middle; inflorescence shortly branched, narrowly paniculate orthyrsoid ............................................................................................................................................................................... 2. I. glabra1. Isoglossa collina (T. Anderson) B. Hansen, Nordic J. Bot. 5:12. 1985.叉 序 草 cha xu caoJusticia collina T. Anderson, J. Linn. Soc., Bot. 9: 515.1867; Chingiacanthus patulus Handel-Mazzetti; Dianthera collina(T. Anderson) C. B. Clarke; D. sinensis W. W. Smith.Herbs 40–100 cm tall. Stems prostrate and rooting at basebut apically erect, terete, slender, slightly puberulent whenyoung, later glabrescent. Petiole 1–3 cm, brownish pubescent;leaf blade ovate to ovate-elliptic, 3.5–11 × 2–4.8 cm, abaxiallysparsely puberulent especially along veins, adaxially glabrous,secondary veins 5–7 on each side of midvein, base cuneate oron uppermost leaves rounded, margin ± entire, apex acuminate.Inflorescences terminal or axillary from upper leaf axils, panicles,5–18 × 3–10 cm; rachis gland-tipped puberulent or subgla-
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ACANTHACEAE 375lary or terminal, 2-
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ACANTHACEAE 377Two species: China,
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ACANTHACEAE 379Vines, herbaceous. S
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ACANTHACEAE 383Type 10. S. tetraspe
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ACANTHACEAE 38525b. Plants strongly
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